Fleshy fruit types

Fruit Types
Fleshy fruit types
Berry
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Drupe
Pome
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Berry
A berry is a simple, indehiscent fruit derived from a single, compound pistil.
The outer pericarp has a distinct outer skin (exocarp) and the flesh (mesocarp)
is usually fleshy.
Kiwi (Actinidia)
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Berry
Depending on the texture of the outer pericarp (skin) and whether the inner fruit
is fleshy or dry, berries can be separated into four distinct groups.
Tomato (Solanum)
Orange (Citrus)
Squash (Cucurbita)
Bacca
Hesperidium
Pepo
Thin outer pericarp;
inner fleshy.
Thick outer pericarp;
inner fleshy with many
segments.
Thick leathery outer
pericarp; inner fleshy
without segments.
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Cannon ball tree
(Couroupita )
Amphisarcum
Thin hard outer
pericarp; inner not
fleshy.
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Berry - Bacca
A bacca is a true berry with fleshy fruit containing many seeds where the entire
pericarp is soft. The pericarp consists of three layers, the outer exocarp (also
called the epicarp), middle fleshy mesocarp (also called sarcocarp), and inner
endocarp. In berries, the endocarp may not be distinct from the mesocarp.
Seed
Mesocarp
Exocarp
Persimmon (Diospyros)
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Berry - Bacca
Grapes produce a cluster of berries.
Seeds
Exocarp
Mesocarp
Grape (Vitis)
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Exocarp
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Berry - Bacca
Although tomatoes are often thought of as a vegetable, the fruits are berries.
Seeds
Mesocarp
Tomato (Solanum)
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Berry - Bacca
Tomatillo is a berry wrapped in a papery husk formed from the calyx (sepals).
Husk
Mesocarp
Exocarp
Seeds
Exocarp
Tomatillo (Physalis)
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Berry - Bacca
Cacti produce berries with many small seeds.
Dragon fruit (Hylocereus )
Cereus
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Mammalaria
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Berry - Hesperidium
A hesperidium is a specialized berry with a leathery outer rind (exocarp).
It is the common fruit type in Citrus.
Orange
Lemon
Citron
(Citrus )
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Berry - Hesperidium
In Citrus, the leathery outer rind
is the exocarp (also flavedo).
Exocarp
Mesocarp
Just under the exocarp is a
spongy middle mesocarp layer
(also albedo).
Segment
The inner edible portion of the
fruit is composed of juice sacs
(vesicles) that are actually hairs
that form on the inner part of
the ovary.
The fruit is septate with each
segment representing a separate
carpel within the ovary.
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Septa
Juice
vesicles
Seeds
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Berry - Hesperidium
There is a mutation in Citrus called ‘Buddha's Hand’ that nicely shows that a
Citrus fruit is composed of fused carpels. The carpels in ‘Buddha’s Hand’ do
not completely fuse resulting in finger-like growths in the fruit each
representing a single carpel or segment.
Fused carpels
Individual carpels
Carpel
Ovary
Ovary
Buddha's Hand
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Carpel
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Berry - Pepo
A pepo is also a specialized berry with a leathery outer covering (exocarp) at
maturity. It is common in the cucurbit family (Cucurbitaceae). It differs from a
hesperidium because it lacks obvious septate carpels.
Exocarp
Mesocarp
Exocarp
Seeds
Winter squash (Cucurbita)
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Berry - Pepo
Pumpkins produce the largest fruits of all plants. In competitions held
around the world, the largest pumpkin weighed in at over 1,500 pounds.
Pumpkins (Cucurbita)
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Berry - Pepo
Pepos can also be found outside the cucurbit family and
include papaya, passion fruit and banana.
Papaya (Carica)
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Passion fruit (Passiflora)
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Banana (Musa)
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Berry - Amphisarcum
An amphisarcum is a relatively new term
to describe fleshy indehiscent fruits
that are berry-like, but have a dry outer
pericarp layer and firm inner flesh.
Gourds are a good example of this type
of fruit.
Gourd (Lagenaria)
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Berry - Amphisarcum
Other plants with this fruit type include the cannonball tree, sausage tree, and
baobab. These large fruits tend to open as they crash to the ground under their
own weight.
Cannonball tree (Couroupita )
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Sausage tree (Kigelia )
Baobab (Adansonia)
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Drupe
A drupe is usually a single-seeded, fleshy fruit with a thin outer skin (exocarp or
epicarp), a soft fleshy mesocarp and a hard endocarp. In members of the rose
family (Rosaceae) like cherry and plum, the endocarp is called a stone.
Exocarp
Mesocarp
Endocarp
Apricot (Prunus)
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Drupe
Inside the stony endocarp is the seed. The endocarp takes over the protective
function usually performed by the seed coat.
Endocarp
Seed
Plum (Prunus)
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Drupe
Dogwood fruit is a drupe. The exocarp is bright red, mesocarp fleshy and the
endocarp hard and stony. In nature, the drupe’s soft parts decompose, but the
endocarp remains and the true seed must germinate through the hard endocarp.
Flowering dogwood (Cornus)
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Drupe
Drupes are a common fruit type that occurs across diverse plant families. Olive is in
the Oleaceae, pepper in the Piperaceae, and mango in the Anacardiaceae.
Pepper (Piper)
Olive (Olea)
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Mango (Mangifera)
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Drupe
Usually a drupe contains only a single seed. However, there are example where
there are numerous seeds within a drupe as occurs in holly. In these cases, the
single, nut-like seeds in the drupe are called pyrenes.
Pyrenes
Holly (Ilex) produces three pyrenes per drupe.
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Drupe
Coffee fruits (often referred to as
“cherries”) are botanically drupes
containing two pyrenes (“beans”).
Coffee
arabica
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Drupe
There are a number of drupe-like fruits that vary from the classic
definition of a drupe because their mesocarp is not fleshy.
A newer term used to describe this type of fruit is a nuculanium.
Exocarp
Endocarp
Seed
Almond (Prunus amygdalum)
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Drupe
Coconut is also an example of a nuculanium. In coconut, the exocarp and mesocarp
are resistant to salt water allowing water dispersal of coconut fruits.
Exocarp
Exocarp
Mesocarp
Endocarp
Seed
Seed
coat
Coconut (Cocos nucifera)
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Pome
A pome is a fleshy fruit where the inner fruit and seeds (inside the papery
endocarp) is surrounded by modified floral tube tissue (from the hypanthium) that
grows to cover the botanical fruit. It is found in some members of the rose family
(Rosaceae) like apple, pear and quince. Technically, a pome is also considered an
accessory fruit because it includes floral tissue in fruit development.
Floral tube
tissue
Seeds
Carpel tissue
Remnant
flower tissue
Apple (Malus)
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Pome
Pear shows a similar arrangement of fruit parts to apple.
Floral tube tissue
Seed
Carpel tissue (endocarp)
Remnant flower tissue
Pear (Pyrus)
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Pome
In the partially matured quince fruit you can readily see the that the fruit
develops from the floral tube (hypanthium) formed below the sepals (calyx).
Floral tube tissue
Sepals
Stamens
Flowering quince (Chaenomoles)
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