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William R. Graves
Curriculum Vitae – March, 2015
Synopsis Dr. Graves is an experienced scholar and administrator of an AAU-­‐member, land-­‐grant, research-­‐intensive university. Bill has been a change-­‐agent in his role as the sole associate dean of his graduate college for nearly five years. Numerous policies and processes have been updated, often radically, during his administrative tenure. This has increased efficiency while broadening services to students, faculty, and staff. Bill’s success with fundraising includes procurement of external grants from diverse federal and private sources, as well as in-­‐kind contributions from university partners, including the Boeing Company, which collaborated with Bill’s college on use of lean principles to improve efficiency in graduate programs across campus. Bill has blended administration with teaching, research, and outreach. His scholarship is documented in a book and in more than 100 publications in refereed journals. Bill has received numerous honors, including awards to recognize his promotion of diversity, his service as a student mentor, his outstanding journal publications, and the impacts on industry resulting from his research. EDUCATION
Doctor of Philosophy – Purdue University – 1988
Dissertation: Urban Root-zone Temperatures and Their Impact on Tree Hydrology and Growth
Master of Science – Iowa State University – 1984
Thesis: The Role of Water Stress and Ethylene in Ficus benjamina L. Leaf Abscission
Bachelor of Science – Iowa State University – 1981
PROFESSIONAL EXPERIENCE
Iowa State University; Ames, Iowa
2010 – present
Associate Dean of the Graduate College
1998 – present
Professor, Department of Horticulture and Interdepartmental
Graduate Program in Ecology and Evolutionary Biology
1995 – 1998
Associate Professor
1992 – 1995
Assistant Professor
Stanford University; Stanford, California
2004 – 2005
Visiting Scholar, Department of Biological Sciences
University of Delaware; Lewes, Delaware
2000 – 2001
Visiting Scientist, Graduate College of Marine Studies
University of Maryland; College Park, Maryland
1988 – 1992
Assistant Professor, Department of Horticulture
2 ADMINISTRATIVE EXPERIENCE AT IOWA STATE UNIVERSITY
Associate Dean of the Graduate College, 2010 - present.
Dr. Graves oversees all functions of the college, which serves nearly
5,000 graduate students and confers the doctorate through 80 majors,
master’s degrees in 108 majors, and certificates in 45 programs.
Primary duties and activities include:
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direct or indirect supervision of 15 to 20 employees
oversee and manage budget of approximately $5 million
strategically plan in conjunction with the dean and assistant dean
manage foundation funds donated to support graduate education
advise the elected Graduate Council about academic policies
monitor policy adherence by students, and resolve waiver requests
contribute to recruiting underrepresented students, a college priority
assist programs for underrepresented students (McNair, etc.)
grant final approval for admission, and, subsequently, for graduation
ensure reporting of data to federal agencies and organizations
with dean, lead meetings of program directors of graduate education
co-lead meetings with graduate-support personnel from programs
process appointments of students to assistantships and fellowships
process appointments of postdoctoral research associates
assist students who are in conflict with faculty mentors
oversee thesis/dissertation formatting checks
advise graduate student and postdoctoral associations
seek external funds and other forms of program support
oversee college’s Academic Communication Program
evaluate and allocate funds to interdepartmental programs
advise faculty who develop new degree and certificate programs
monitor students on academic probation
approve removal from restricted or provisional admission status
oversee awards programs for students and graduate faculty
Selected examples of major changes in graduate education fostered by Associate
Dean Graves, 2010 - present.
new electronic process to develop students’ committees and course plans
new budgeting model for interdepartmental graduate programs
new Academic Communications Program for graduate students
new policies for students on residency, use of undergraduate courses, and
distance participation in oral examinations
• new services for career exploration and postdoctoral scholars
• new, paperless environment for student records
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3 Director of Graduate Education — interdisciplinary graduate studies major, 2010 present.
The director of graduate education is responsible for overseeing the
development and implementation of policies governing master’s programs in
this major. Because the major can be pursued in collaboration with various
disciplines, the director must coordinate communication to ensure an
outstanding experience for students and faculty who participate in the major.
Director of Graduate Education — horticulture major, 1998 - 2010.
The director of graduate education is responsible for overseeing the
development and implementation of policies governing Ph.D. and M.S.
programs in horticulture. Through interaction with the departmental graduate
education committee, the director helps to recruit students to the major and
makes recommendations on admissions. The director also is responsible for
facilitating dialogue about the program among students and members of the
graduate faculty. The director facilitates communication between the
graduate college and the graduate faculty and students in horticulture.
Program Reviewer — Department of Horticultural Science, University of
Minnesota, October, 2004.
Dr. Graves was one of five members of a review team chosen from a national
pool to review the scholarly programs in horticultural science. He focused on
research and graduate education.
Program Reviewer — Department of Horticulture, University of Georgia, October
and November, 2002.
Academic programs in horticulture were reviewed by Dr. Graves and other
members of a peer group chosen from a national pool.
Strategic-planning Panelist and Topic Expert — National Urban and Community
Forestry Research and Technology Transfer Summit, December, 2002, Chicago,
Illinois.
Dr. Graves was one of several experts chosen from across the United States
to participate in strategic planning and in development of the National Urban
and Community Forestry Action Plan for the next decade.
4 Program Reviewer — Department of Horticulture, University of Arkansas,
November, 2001.
Dr. Graves was a member of a review team chosen from a national pool to
review the academic programs in horticulture.
Program Reviewer — Department of Horticulture, Cornell University, October,
2000.
The graduate program was Dr. Graves’ area of emphasis during a review of
horticultural academics by a five-member team selected from across the
United States.
Elected Member of Graduate Council — Agriculture and Life Sciences Division,
Iowa State University, 1996 - 1999.
The graduate council sets campus-wide policy regarding rules and procedures
of the graduate college. Dr. Graves participated on a subcommittee that
reviewed the status of thesis and non-thesis graduate degree programs and
Iowa State. He also led a subcommittee on establishing a preparing future
faculty program on campus.
PROFESSIONAL DEVELOPMENT RELATED TO ADMINISTRATION
Council of Graduate Schools – Conference Participation
2010 Annual Meeting; Washington, D.C.
2011 New Deans Institute and Summer Workshop; Monterey, California
2011 Annual Meeting; Scottsdale, Arizona
2012 Summer Workshop; Boston, Massachusetts
2012 Annual Meeting; Washington, D.C.
2013 Summer Workshop; San Juan, Puerto Rico
2013 Annual Meeting; San Diego, California 2014
Summer Workshop;Portland,Oregon
2014 Annual Meeting; Washington, D.C.
Activities with Midwestern Association of Graduate Schools
2011 Annual Meeting; Milwaukee, Wisconsin
leader of hot-topics discussion session on distance education
2012 Annual Meeting; Chicago, Illinois
5 2012 – 2014: Member of Outstanding Master’s Thesis Selection
Committee
2013 Annual Meeting; Minneapolis, Minnesota
2014 Annual Meeting; Chicago, Illinois
co-presenter of session on lean-process collaboration between Iowa
State’s Graduate College and the Boeing Company
Nominated in 2015 to serve on Executive Committee
Chosen as one of 50 deans nationally to participant in NSF-funded Graduate
Deans Workshop on Institutionalizing Interdisciplinary Graduate Education;
Arlington, Virginia; 2-3 April 2012
COMMITTEE SERVICE RELATED TO ADMINISTRATION
• Elected or Appointed Member, Department Head's/Chair’s Advisory Committee
— Department of Horticulture, Iowa State University, various years 1993 present
• Chair of Graduate Education Committee — Department of Horticulture, Iowa
State University, various years 1998 - 2010
• Promotion and Tenure Committee — College of Agriculture, Iowa State
University, 1998 - 2001
• Promotion and Tenure Committee — Department of Horticulture, various years
(service as chair during some years) 1999 - present
• Preparing Future Faculty Advisory Committee — Graduate College, Iowa State
University, 2001 - 2002
6 SERVICE FOR GRADUATE STUDENTS AND OTHER SCHOLARS
Listed in reverse chronological order of matriculation or graduation date are all graduate
students being advised by Dr. Graves, and students who have completed a graduate
degree program with Dr. Graves as major professor or co-major professor. Most
students were competitively recruited from other institutions and were placed in
excellent jobs upon degree completion. Dr. Graves’ doctoral advisees have taken
tenure-track positions at the University of Illinois, the University of Florida, New Mexico
State University, Utah State University, the University of Nevada, and the University of
Maine. Other graduates of his program accepted employment offers at North Carolina
State University, the University of Arizona, the University of Kentucky, the University of
Wisconsin, Iowa State University, the Natural Resources Conservation Service of the
United States Department of Agriculture, the Des Moines Botanical Garden, the
Environmental Protection Agency, Monsanto Corporation, and other private companies.
Ph.D. students
M.S. students
Zachary Hudson (current)
Kalynn Doebel (current)
Bryan J. Peterson (2013)
Frank Balestri (current)
Olivia Lenahan
Kenneth McCabe
(2009)
(current) Nickolee Zollinger
(2009)
Kelly D. Norris
(2012)
J. Ryan Stewart (2006)
Bryan J. Peterson (2009)
Heidi A. Kratsch (2004)
Matthew S. Helgeson (2009)
Annita G. Peterson (2004)
M. Tiffany Laws (2004)
James. A. Schrader (2002)
Sarah J. Gardner (2001)
Carol M. Foster (1998)
Michael L. Adam
Rolston St. Hilaire (1998)
James A. Schrader (1999)
Kimberly Klock
James A. Zwack (1997)
(1995)
Hongyi Zhang
(2000)
(1994)
J. Giridhar B. Pai (1994)
Lorna C. Wilkins
Lenore J. Nash
(University of Maryland, 1992)
Janet C.M. Batzli
(University of Maryland, 1993)
(University of Maryland, 1991)
7 In addition, Dr. Graves has served as:
• mentor for two postdoctoral associates;
• faculty advisor for the Graduate Student Horticulture Society (student club); and
• member of numerous program of study committees, including several for students
in colleges other than his own.
REFEREED PUBLICATIONS
Peterson, B.J. and W.R. Graves. 2015. Chloroplast phylogeography of Dirca palustris L.
indicates populations near the glacial boundary at the Last Glacial Maximum in eastern
North America. Biogeography: accepted pending revision.
Madbouly, S., S. Yang, M. Kessler, D. Grewell, J. Schrader, and W.R. Graves. 2015.
Processing and characterization of bio-based poly(hydroxyalkanoate) (PHA)/poly(amide)
(PA) blends: Improved flexibility and impact resistance of PHA-based plastics. Journal of
Applied Polymer Science: in press.
Schrader, J.A., K.G. McCabe, G. Srinivasan, K. Haubrich, D. Grewell, S. Madbouly, and
W.R. Graves. 2015. Development and evaluation of bioplastic containers for sustainable
greenhouse and nursery production. Acta Horticulturae: in press.
Yang, S., S.A. Madbouly, J.A. Schrader, G. Srinivasan, D. Grewell, K.G. McCabe, M.R.
Kessler, and W.R. Graves. 2015. Characterization and biodegradation behavior of biobased poly(lactic acid) and soy protein blends for sustainable horticultural applications.
Green Chemistry 17:380-393.
Lu, H., S.A. Madbouly, J.A. Schrader, G. Srinivasan, K.G. McCabe, D. Grewell, M.R.
Kessler, and W.R. Graves. 2014. Biodegradation behavior of poly (lactic acid)
(PLA)/distiller's dried grains with soluble (DDGS) composites. ACS Sustainable
Chemistry & Engineering 2:2699-2706.
Lu, H., S.A. Madbouly, J.A. Schrader, M.R. Kessler, D. Grewell, and W.R. Graves. 2014.
Novel bio-based composites of polyhydroxyalkanoate (PHA)/distillers dried grains with
solubles (DDGS). RSC Advances 4:39802-39808.
McCabe, K.G., J.A. Schrader, S. Madbouly, D. Grewell, and W.R. Graves. 2014.
Evaluation of biopolymer-coated fiber containers for container-grown plants.
HortTechnology 24:439-448.
Madbouly, S.A., J.A. Schrader, G. Srinivasan, K. Liu, K.G. McCabe, D. Grewell, W.R.
Graves, and M.R. Kessler. 2014. Biodegradation behavior of bacterial-based
polyhydroxyalkanoate (PHA) and DDGS composites. Green Chemistry 16:1911-1920.
Grewell, D., G. Srinivasan, J. Schrader, W. Graves, and M. Kessler. 2014. Sustainable
materials for horticultural application. Plastics Engineering 70(3):44-52.
8 Schrader, J.A., G. Srinivasan, D. Grewell, K.G. McCabe, and W.R. Graves. 2013.
Fertilizer effects of soy-plastic containers during crop production and transplant
establishment. HortScience 48:724-731.
Peterson, B.J. and W.R. Graves. 2013. Responses to root-zone water content of shrub
congeners from eastern North America and Mediterranean California. HortScience 48:
715-719.
Norris, K.D. and W.R. Graves. 2012. A narrowly endemic Dirca from Mexico outperforms
its broadly distributed congener in the Upper Midwest. HortScience 47:1445-1448.
Schrader, J.A. and W.R. Graves. 2011. Taxonomy of Leitneria (Simaroubaceae)
resolved by ISSR, ITS, and morphometric characterization. Castanea 76:313-338.
Peterson, B.J. and W.R. Graves. 2011. Reproductive ecology of Dirca palustris L.
(Thymelaeaceae). Castanea 76:237-244.
Graves, W.R. and A.L. Thomas. 2011. Survival and growth of Callicarpa americana of
northern and southern origin in U.S.D.A. hardiness zones 5 and 6. Journal of
Environmental Horticulture 29:9-13.
Peterson, B.J., W.R. Graves, and J. Sharma. 2011. Phenotypic and genotypic diversity
of eastern leatherwood in five populations that span its geographic distribution. American
Midland Naturalist 165:1-21.
Lenahan, O.M., W.R. Graves, and R. Arora. 2010. Cold hardiness and deacclimation of
Styrax americanus from three provenances. HortScience 45:1819-1823.
Helgeson, M.S., W.R. Graves, D. Grewell, and G. Srinivasan. 2010. Zein-based
bioplastic containers alter root-zone chemistry and growth of geranium. Journal of
Environmental Horticulture 28:74-80.
Peterson, B.J. and W.R. Graves. 2009. Variation in development and response to rootzone pH among seedlings of Dirca palustris (Thymelaeaceae) from three provenances.
HortScience 44:1319-1322.
Boyer, N.Z. and W.R. Graves. 2009. NAA is more effective than IBA for rooting stem
cuttings of two Nyssa spp. Journal of Environmental Horticulture 27:183-187.
Helgeson, M.S., W.R. Graves, D. Grewell, and G. Srinivasan. 2009. Degradation and
nitrogen release of zein-based bioplastic containers. Journal of Environmental
Horticulture 27:123-127.
Peterson, B.J., Graves, W.R., and J. Sharma. 2009. Color of pubescence on bud scales
conflicts with keys for identifying species of Dirca (Thymelaeaceae). Rhodora 111:126130.
9 Graves, W.R. 2008. Habitat and reproduction of Dirca mexicana. Rhodora 110:365-378.
Graves, W.R. and J.A. Schrader. 2008. At the interface of phylogenetics and population
genetics, the phylogeography of Dirca occidentalis (Thymelaeaceae). American Journal
of Botany 95:1454-1465.
Sharma, J., J.A. Schrader, and W.R. Graves. 2008. Ecology and phenotypic variation of
Leitneria floridana (Leitneriaceae) in disjunct native habitats. Castanea 73:94-105.
Schrader, J.A. and W.R. Graves. 2008. Nodulation and growth of Alnus nitida and Alnus
maritima inoculated with species-specific and nonspecific Frankia. Journal of
Environmental Horticulture 26:29-34.
Schrader, J.A., M.A. Kroggel, and W.R. Graves. 2007. Nodulation and nitrogen-fixing
capacity of rhizobial isolates from China in symbiosis with Maackia amurensis. Journal of
Environmental Horticulture 25:47-50.
Schrader, J.A., W.R. Graves, S.A. Rice, and J.P. Gibson. 2006. Differences in shade
tolerance help explain varying success of two sympatric Alnus species. International
Journal of Plant Sciences 167:979-989.
Stewart, J.R. and W.R. Graves. 2006. Photosynthesis, growth, carbon allocation and fruit
load of Frangula caroliniana and Rhamnus cathartica. International Journal of Plant
Sciences 167:1161-1168.
Graves, W.R., J.A. Schrader, and J. Sharma. 2006. Cold hardiness of the rare Dirca
occidentalis: Comparisons to Dirca palustris from disparate provenances. Journal of
Environmental Horticulture 24:169-172.
Stewart, J.R. and W.R. Graves. 2006. Cold hardiness and vernal bud break of Rhamnus
caroliniana and Rhamnus cathartica. Journal of the American Society for Horticultural
Science 131:345-351.
J. Zhu, A. Zhang, K. Park, T. Baker, B. Lang, R. Jurenka, J.J. Obrycki, W.R. Graves, J.A.
Pickett, D. Smiley, K.R. Chauhan, J.A. Klun. 2006. Sex pheromone of the soybean
aphid, Aphis glycines Matsumura, and its potential use in semiochemical-based control.
Environmental Entomology 35:249-257.
Graves, W.R. 2006. Distribution and reproduction of Dirca occidentalis (western
leatherwood). The Four Seasons (East Bay [Berkeley, California] Regional Parks Botanic
Garden) 12(3):54-61.
Kratsch, H.A., R.J. Gladon, and W.R. Graves. 2006. Aeroponics system for control of
root-zone atmosphere. Environmental and Experimental Botany 55:70-76.
Sharma, J. and W.R. Graves. 2005. Propagation of Rhamnus alnifolia and Rhamnus
lanceolata by seeds and cuttings. Journal of Environmental Horticulture 23:86-90.
10 Voegtlin, D.J., R.J. O'Neil, W.R. Graves, D. Lagos, and H.J.S. Yoo. 2005. Potential
winter hosts of the soybean aphid, Aphis glycines Matsumura. Annals of the
Entomological Society of America 98:690-693.
Yoo, H.J.S., R.J. O'Neil, D.J. Voegtlin, and W.R. Graves. 2005. Host plant suitability of
Rhamnaceae for the soybean aphid, Aphis glycines Matsumura (Homoptera: Aphididae).
Annals of the Entomological Society of America 98:926-930.
Schrader, J.A. and W.R. Graves. 2005. Seed germination of Dirca (leatherwood):
Pretreatments and interspecific comparisons. HortScience 40:1838-1842.
Laws, M.T. and W.R. Graves. 2005. Nitrogen inhibits nodulation and reversibly
suppresses nitrogen fixation in nodules of Alnus maritima. Journal of the American
Society for Horticultural Science 130:496-499.
Stewart, J.R. and W.R. Graves. 2005. Seed germination of Rhamnus caroliniana:
implications for ecology and horticulture. HortScience 40:767-770.
Kratsch, H.A. and W.R. Graves. 2005. Oxygen concentration affects nodule anatomy
and nitrogenase activity of Alnus maritima. Plant, Cell and Environment 28:688-696.
Schrader, J.A., S.J. Gardner, and W.R. Graves. 2005. Resistance to water stress of
Alnus maritima: Intraspecific variation and comparisons to other alders. Environmental
and Experimental Botany 53:281-298.
Hilgers, K.R., C.L. Haynes, and W.R. Graves. 2005. Chemical height control of
containerized seashore mallow. HortTechnology 15:330-332.
Kratsch, H.A. and W.R. Graves. 2004. Location and anatomy of nodules on Alnus
maritima subjected to flooding. Journal of the American Society for Horticultural Science
129:775-780.
Graves, W.R. 2004. Confirmation that Dirca spp. (Thymelaeaceae) reproduce from
rhizomes. Rhodora 106:291-294.
Sharma, J. and W.R. Graves. 2004. Germination of Leitneria floridana seeds from
disjunct populations. HortScience 39:1695-1699.
Stewart, J.R. and W.R. Graves. 2004. Photosynthesis and growth of Rhamnus
caroliniana during drought and flooding: comparisons to the invasive Rhamnus
cathartica. HortScience 39:1278-1282.
Schrader, J.A. and W.R. Graves. 2004. Systematics of Dirca (Thymelaeaceae) based on
ITS sequences and ISSR polymorphisms. Sida 21:517-530.
Kratsch, H.A. and W.R. Graves. 2004. Nitrogen fixation as a stress-avoidance strategy
among actinorhizal (non-legume) trees and shrubs. Journal of Crop Improvement
10:281-304.
11 Sharma, J. and W.R. Graves. 2004. Midwinter cold hardiness of Leitneria floridana from
three provenances. Journal of Environmental Horticulture 22:88-92.
Graves, W.R. and J.A. Schrader. 2004. ‘September Sun’ seaside alder, an autumnblooming shrub native to North America. HortScience 39:438-439.
Voegtlin, D., R.J. O'Neil, and W.R. Graves. 2004. Tests of suitability of overwintering
hosts of Aphis glycines: identification of a new host association with Rhamnus alnifolia
L’Héritier. Annals of the Entomological Society of America 97:233-234.
Schrader, J.A. and W.R. Graves. 2004. Systematics of Alnus maritima (Seaside Alder)
resolved by ISSR polymorphisms and morphological characters. Journal of the American
Society for Horticultural Science 129:231-236.
Haynes, C.L. and W.R. Graves. 2004. Kosteletzkya virginica can be rooted from leafy or
leafless stem cuttings. Journal of Environmental Horticulture 22:173-175.
Stewart, J.R. and W.R. Graves. 2004. Photosynthesis and growth of Carolina buckthorn
(Rhamnus caroliniana Walter) during drought and flooding: comparisons to the invasive
common buckthorn (Rhamnus cathartica L.). Acta Horticulturae 630, ISHS 2004:143146.
Kratsch, H.A. and W.R. Graves. 2004. Ultrastructure of nodules from Alnus maritima.
Acta Horticulturae 630, ISHS 2004:231-236.
Schrader, J.A. and W.R. Graves. 2004. Ecophysiology, phenology, and depth of cold
acclimation in the three subspecies of Alnus maritima. Acta Horticulturae 630, ISHS
2004:199-205.
Graves, W.R. 2004. Adaptation evaluated: case studies of underused trees and shrubs.
Acta Horticulturae 630, ISHS 2004:125-135.
Graves, W.R. and J.L. Gallagher. 2003. Resistance to salinity of Alnus maritima from
disjunct wetlands: symptoms of salt injury, comparison to other shrubs, and effect of
inundation. Wetlands 23:394-405.
Schrader, J.A. and W.R. Graves. 2003. Phenology and depth of cold acclimation in the
three subspecies of Alnus maritima. Journal of the American Society for Horticultural
Science 128:330-336.
Adam, M.L., J.M. Kelly, W.R. Graves, and P. Dixon. 2003. Net nitrate uptake by Acer
rubrum is a function of root-zone temperature. Journal of Plant Nutrition 26:203-222.
Schrader, J.A. and W.R. Graves. 2002. Infraspecific systematics of Alnus maritima
(Betulaceae) from three widely disjunct provenances. Castanea 67:380-401.
Gladon, R.J., W.R. Graves, and J.M. Kelly. 2002. Publishing in plant science journals: A
course that jump-starts professional careers. Journal of Natural Resources and Life
Sciences Education 31:88-93.
12 Graves, W.R. 2002. IBA, juvenility, and position on ortets influence propagation of
Carolina buckthorn from softwood cuttings. Journal of Environmental Horticulture 20: 5761.
Graves, W.R., M.A. Kroggel, and M.P. Widrlechner. 2002. Photosynthesis and shoot
health of five birch and four alder taxa after drought and flooding. Journal of
Environmental Horticulture 20:36-40.
St. Hilaire, R., W.R. Graves, and R.L. Small. 2001. Variation in Taq1-digested DNA of
sugar and black maples is independent of taxon and geography. HortScience 36:13271328.
St. Hilaire, R. and W.R. Graves. 2001. Stability of provenance differences during
development of hard maple seedlings irrigated at two frequencies. HortScience 36:654657.
Foster, C.M., H.T. Horner, and W.R. Graves. 2000. Accumulation of ENOD2-like
transcripts in non-nodulating woody papilinoid legumes. Plant Physiology 124:741-750.
Kelly, J.M., W.R. Graves, and A.S. Aiello. 2000. Nitrate uptake kinetics for rooted
cuttings of Acer rubrum L. Plant and Soil 221:221-230.
Schrader, J.A. and W.R. Graves. 2000. Alnus maritima: a rare woody species from the
New World. The New Plantsman 7:74-82.
Schrader, J.A. and W.R. Graves. 2000. Propagation of Alnus maritima from softwood
cuttings. HortScience 35:293-295.
Schrader, J.A. and W.R. Graves. 2000. Timing of seed dispersal may limit the
reproductive success of Alnus maritima. Castanea 65:69-77.
Schrader, J.A. and W.R. Graves. 2000. Seed germination and seedling growth of Alnus
maritima from its three disjunct populations. Journal of the American Society for
Horticultural Science 125:128-134.
St. Hilaire, R. and W.R. Graves. 1999. Foliar traits of sugar maples and black maples
near 43 °N latitude in the eastern and central United States. Journal of the American
Society for Horticultural Science 124:605-611.
Graves W.R., C.M. Foster, F.M. Rosin, and J.A. Schrader. 1999. Two early nodulation
genes are not markers for the capacity of leguminous nursery crops to form root nodules.
Journal of Environmental Horticulture 17:126-129.
Dosmann, M.S., J.K. Iles, and W.R. Graves. 1999. Drought avoidance in katsura by
drought-induced leaf abscission and rapid refoliation. HortScience 34:871-874.
Zwack, J.A., W.R. Graves, and A.M. Townsend. 1999. Variation among red and
Freeman maples in response to drought and flooding. HortScience 34:664-668.
Foster, C.M., H.T. Horner, and W.R. Graves. 1998. Nodulation response of woody
Papilinoid species after inoculation with rhizobia and soils from Hawaii, Asia, and North
America. Plant and Soil 205:103-111.
13 Iles, J.K., W.R. Graves, A.S. Aiello, C.L. Haynes, and K.E. Stone. 1998. Midwestern
retail garden centers identify satisfaction/dissatisfaction with supplying wholesale
nurseries. Journal of Environmental Horticulture 16:243-247.
Aiello, A.S. and W.R. Graves. 1998. Success varies when using subirrigation instead of
mist to root softwood cuttings of woody taxa. Journal of Environmental Horticulture
16:42-47.
Foster, C.M., H.T. Horner, and W.R. Graves. 1998. Isolation of a 14-3-3 brain protein
homolog from nodules of Maackia amurensis Rupr. & Maxim. (accession no. AF039709)
(PGR 98-061) Plant Physiology 116:1604.
Foster, C.M., W.R. Graves, and H.T. Horner. 1998. ENOD2 cDNA clone from nodules of
Maackia amurensis Rupr. & Maxim. (accession no. AF039708) (PGR 98-060). Plant
Physiology 116:1604.
Zwack, J.A., W.R. Graves, and A.M. Townsend. 1998. Leaf water relations and plant
development of three Freeman maple cultivars subjected to drought. Journal of the
American Society for Horticultural Science 123:371-375.
Aiello, A.S. and W.R. Graves. 1997. Container medium and nitrogen form affect
production of Amur maackia. HortScience 32:1200-1203.
Geater, C.A., G.R. Nonnecke, W.R. Graves, A.S. Aiello, and C.A. Dilley. 1997. High rootzone temperatures inhibit growth and development of Fragaria species. Fruit Varieties
Journal 51:94-101.
Graves, W.R. and A.S. Aiello. 1997. Root-zone temperatures above 29 °C reduce
transpiration, water potential, and growth of silver maple indigenous to Minnesota and
Mississippi. Journal of the American Society for Horticultural Science 122:195-199.
Klock, K.A., H.G. Taber, and W.R. Graves. 1997. Root respiration and phosphorus
nutrition of tomato plants grown at 36 °C root-zone temperature. Journal of the American
Society for Horticultural Science 122:175-178.
Zhang, H., W.R. Graves, and A.M. Townsend. 1997. Water loss and survival of stem
cuttings of two maple cultivars held in subirrigated rooting medium at 24 to 33 °C.
HortScience 32:129-131.
Graves, W.R. and H. Zhang. 1996. Relative water content and rooting of subirrigated
stem cuttings in four environments without mist. HortScience 31:866-868.
Aiello, A.S. and W.R. Graves. 1996. Two leguminous tree species differ in growth and
ion uptake in Hoagland solution: Phosphorus toxicity in Amur maackia. Journal of Plant
Nutrition 19:1061-1073.
Klock, K.A., W.R. Graves, and H.G. Taber. 1996. Growth and phosphorus, zinc, and
manganese content of tomato, muskmelon, and honey locust at high root-zone
temperatures. Journal of Plant Nutrition 19:795-806.
14 Pai, J.G.B. and W.R. Graves. 1995. Seed source affects seedling development and
nitrogen fixation of Maackia amurensis. Journal of Environmental Horticulture 13:142146.
Wilkins, L.C., W.R. Graves, and A.M. Townsend. 1995. Responses to high root-zone
temperature among cultivars of red maple and Freeman maple. Journal of Environmental
Horticulture 13:82-85.
Pai, J.G.B. and W.R. Graves. 1995. Influence of inoculant form and applied nitrogen on
growth and root nodulation of Maackia amurensis. Journal of Environmental Horticulture
13:40-42.
Zhang, H. and W.R. Graves. 1995. Subirrigation to root stem cuttings: Comparison to
intermittent mist and influence of fertilization. HortTechnology 5:265-268.
Wilkins, L.C., W.R. Graves, and A.M. Townsend. 1995. Development of plants from
single-node cuttings differs among cultivars of red maple and Freeman maple.
HortScience 30:360-362.
Graves, W.R., S.R. Anfinson, and K.K. Lappegard. 1995. Leaching fraction and
fertilization effects on growth of three woody legumes inoculated with rhizobia.
HortScience 30:72-73.
Graves, W.R. 1994. Development of seedlings of sugar maple and black maple irrigated
at various frequencies. HortScience 29:1292-1294.
McNamara, S., W.R. Graves, and H. Pellett. 1994. Propagation of Maackia amurensis
from softwood cuttings. Journal of Environmental Horticulture 12:147-149.
Behboudian, M.H., W.R. Graves, C.S. Walsh, and R.F. Korcak. 1994. Water relations,
mineral nutrition, growth, and 13C discrimination in two apple cultivars under daily
episodes of high root-medium temperature. Plant and Soil 162:125-133.
Graves, W.R. and S.R. Anfinson. 1994. Influence of reduced leaching on growth of
seedlings of black locust inoculated with rhizobia. Journal of the Iowa Academy of
Science 101:54-55.
Graves, W.R. 1994. Urban root-zone temperatures and their potential impact on trees.
Journal of Arboriculture 20:24-27.
Nash, L.J. and W.R. Graves. 1993. Drought and flood stress effects on plant
development and leaf water relations of five tree species native to bottomland habitats.
Journal of the American Society for Horticultural Science 118:845-850.
Graves, W.R. and L.C. Wilkins. 1993. An exercise to illustrate biological nitrogen fixation
to plant science students. HortTechnology 3:462-463.
Batzli, J.M., W.R. Graves, and P. van Berkum. 1992. Diversity among rhizobia effective
on Robinia pseudoacacia L. Applied and Environmental Microbiology 58:2137-2143.
Graves, W.R. 1992. Influence of hydroponic culture method on morphology and hydraulic
conductivity of roots of honey locust. Tree Physiology 11:205-211.
15 Graves, W.R. and W. van de Poll. 1992. Further evidence that American yellowwood
does not form nitrogen-fixing symbioses with rhizobia. HortScience 27:1137.
Batzli, J.M., W.R. Graves, and P. van Berkum. 1992. Isolation and characterization of
rhizobia effective with Maackia amurensis. Journal of the American Society for
Horticultural Science 117:612-616.
Graves, W.R. 1991. Growth and iron content of three legume tree species at high rootzone temperature. Journal of Arboriculture 17:313-317.
Graves, W.R. and L.C. Wilkins. 1991. Growth of honey locust seedlings during high rootzone temperature and osmotic stress. HortScience 26:1312-1315.
Graves, W.R., R.J. Joly, and M.N. Dana. 1991. Water use and growth of honey locust
and tree-of-heaven at high root-zone temperature. HortScience 26:1309-1312.
Graves, W.R. 1990. Stratification not required for tree-of-heaven seed germination. Tree
Planters' Notes 41(2):10-12.
Graves, W.R., M.N. Dana, and R.J. Joly. 1989. Influence of root-zone temperature on
growth of Ailanthus altissima (Mill.) Swingle. Journal of Environmental Horticulture 7:7982.
Graves, W.R., M.N. Dana, and R.J. Joly. 1989. Root-zone temperature affects water
status and growth of red maple. Journal of the American Society for Horticultural Science
114:406-410.
Graves, W.R. and M.N. Dana. 1987. A system to control root-zone temperature.
HortScience 22:957-958.
Graves, W.R. and M.N. Dana. 1987. Root-zone temperature monitored at urban sites.
HortScience 22:613-614.
Graves, W.R. and R.J. Gladon. 1985. Water stress, endogenous ethylene, and Ficus
benjamina L. leaf abscission. HortScience 20:273-275.
BOOKS
Gladon, R.J., W.R. Graves, and J.M. Kelly. 2011. Getting Published in Life Sciences.
Wiley-Blackwell; Hoboken, NJ; 356 pages.
16 SUMMARY OF PUBLISHED ABSTRACTS AND PRESENTATIONS
Career total: 128
Selected conference venues:
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Ecological Society of America
American Society for Horticultural Science
International Plant Propagator’s Society
Plant Biology
Botanical Society of America
California Native Plant Society
Iowa State University Shade Tree Short Course
American Association for the Advancement of Science
North American Nitrogen Fixation Conference
Midwestern Rare Plant Conference
Ohio Nursery Short Course
XXII International Congress of Entomology
International Society of Arboriculture
Metropolitan Tree Improvement Alliance
Arboriculture and Urban Forestry Symposium
Most Recent Published Abstracts (all at conference of American Society for Horticultural
Science; Orlando, Florida; July, 2014)
F.D. Balestri and W.R. Graves. Pretreatment Effects on Seed Germination: An
Interspecific Comparison of Two Nyssa Species. (poster; graduate student
presented)
B.J. Peterson and W.R. Graves. Faster and Cheaper Microsatellite Projects with
Plants. (oral; former graduate student [now assistant professor] presented)
J.D. Mahoney, F.D. Balestri, and W.R. Graves. Iowa State University Heritage
Tree Program: Preserving the Genetic Heritage of Iowa State’s Award-winning
Campus. (poster; undergraduate student presented)
C.J. Currey, K. McCabe, J. Schrader, G. Srinivasan, D. Grewell, and W.R.
Graves. Soy-based Biocontainers Allow for Reduced Fertilizer Inputs. (oral;
faculty colleague presented)
J. Schrader, K. McCabe, G. Srinivasan, D. Grewell, K. Haubrich, Samy
Madbouly, H. Kratsch, C. Currey, and W.R. Graves. Evaluation and Classification
of Bioplastic Plant Containers. (oral; Graves presented)
17 INSTRUCTION
Dr. Graves co-developed and annually co-instructed for nearly 20 years a graduate
course, Publishing in Biological Science Journals (Agron/Hort/NREM 529; 2 or 3
credits). In 2014, this course was used as a model for a new class that was offered to
graduate students in all disciplines at Iowa State. The new course, offered from the
Graduate College rather than from one or more departments, is titled Preparing
Publishable Thesis Chapters (GR ST [Graduate Studies] 529). Each student in this
course uses an innovative, practical approach for preparing a manuscript for submission
to a refereed journal in her or his discipline. In addition, students are sensitized to the
linguistic conventions that differ among scholars in various fields of inquiry and to the
importance of issues ranging from formatting guidelines to avoidance of plagiarism.
Before assuming the role of associate dean, Dr. Graves was responsible for Hort 530,
Research Orientation (2 credits each fall semester), the introductory core course for
students in the horticulture graduate major. He also contributed to Hort 698,
Horticulture Teaching Practicum (1-credit each spring semester), which allows graduate
students to gain instructional skills and to be exposed to critical issues in higher
education. Dr. Graves has been responsible for Hort 610, Graduate Seminar, and he
regularly advises with students registered for Hort 699, Thesis and Dissertation
Research. Dr. Graves led EEB (Ecology and Evolutionary Biology) 585, Extended Field
Trip, during spring, 2011. This experiential course was designed around a trip during
spring break of that semester to Stanford University’s Jasper Ridge Biological Preserve
and other venues pertaining to ecological science in the Bay Area of California.
Since joining the faculty at Iowa State in 1992, Dr. Graves has taught at least one
undergraduate course per year on the identification and use of trees, shrubs, and
woody vines. This responsibility continues while serving as associate dean.
AWARDS AND OTHER INDICATORS OF ACCOMPLISHMENT
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Margaret Ellen White Graduate Faculty Award, Iowa State University, 2010
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American Society for Horticultural Science Ornamentals/Landscape & Turf
Distinguished Achievement Award, August, 2005
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Diversity Enhancement Award, College of Agriculture, Iowa State University,
2004
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Norman Jay Colman National Research Award, American Nursery and
Landscape Association, 1999
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Windler Award for Best Systematics Research Paper (with graduate student)
Career Total Invited Presentations of Scholarship: 35
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International projects in China, India, and Mexico resulting in refereed articles
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Study Abroad Course and Trip Planner/Leader/Participation: Italy,
England, South Africa
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Offered but declined Bailey endowed-chair professorship at University of
Minnesota, 1998
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Elected to Graduate Council, Iowa State University
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Elected Member of Osborn Club, Iowa State University
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Associate Editorship – American Society for Horticultural Science
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Editorial Board Member – Journal of Environmental Horticulture
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Grant Review Panel (2002 – 2007) – American Nursery and Landscape
Association
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Research Committee – Landscape Plant Development Center
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Member of Publications, Research, and Paper-award Committees, American
Society for Horticultural Science
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Plant Patent for Alnus maritima subsp. oklahomensis ‘September Sun’ (United
States Plant Patent Number PP18,101, October, 2007)
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Visiting scientist/scholar at University of Delaware and Stanford University
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Doctoral students and postdoctoral associates assumed professional academic
positions at: North Carolina State, Illinois, Florida, Brigham Young, New Mexico
State, Utah State, Nevada, Maine, and Texas Tech
FUNDING SUMMARY
Total (intramural/extramural) secured for scholarship, 2000 – 2015: $3,595,456
Recent extramural funding sources:
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USDA-ARS-Specialty Crops Research Initiative Competitive Grants Program
Horticultural Research Institute
California Academy of Sciences
International Plant Propagators’ Society
California Native Plant Society
J. Frank Schmidt Family Charitable Foundation
Iowa Nursery and Landscape Association Research Foundation
USDA-CSREES Integrated Pest Management Program
Landscape Plant Development Center
United Soybean Board
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