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A Field Guide
To the Permanent Sample Plots
in the Conventional Logging Blocks 28 & 29
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
A Field Guide
To the Permanent Sample Plots
in the Conventional Logging Blocks 28 & 29
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
National Library of Indonesia Cataloging-in-Publication Data
Kartawinata, Kuswata et al.
A field guide to the permanent sample plots in the conventional logging blocks 28 & 29 at CIFOR
Malinau research forest East Kalimantan/by Kuswata Kartawinata, Hari Priyadi, Douglas Sheil,
Soedarsono Riswan, Plinio Sist, Machfudh. Bogor, Indonesia: Center for International Forestry Research
(CIFOR), 2006.
113p.
ISBN: 979-24-4631-1
CAB Int. thesaurus: 1. logging 2. sample plot technique 3. stand density 4. forest trees 5. identification
6. Indonesia 7. guidelines
© 2006 by CIFOR and ITTO
All rights reserved. Published in 2006
Printed by Intiprima Karya
Cover photos by Hari Priyadi and Ahmad Zakaria
Cover design by Eko Prianto
Layout by Catur Wahyu
Maps by Mohammad Agus Salim
Published by
Center for International Forestry Research
Mailing address: P.O. Box 6596 JKPWB, Jakarta 10065, Indonesia
Office address: Jl. CIFOR, Situ Gede, Sindang Barang,
Bogor Barat 16680, Indonesia
Tel.: +62 (251) 622622; Fax: +62 (251) 622100
E-mail: [email protected]
Web site: http://www.cifor.cgiar.org
3
Contents
Foreword
4
Introduction
5
The Plot
8
Access to a Plot
9
Plant Identification
10
Distribution of PSPS
11
Plot# CNV01
12
Plot# CNV02
19
Plot# CNV03
28
Plot# CNV04
35
Plot# CNV05
42
Plot# CNV06
52
Plot# CNV07
60
Plot# CNV08
67
Plot# CNV09
77
Plot# CNV10
87
Plot# CNV11
96
Plot# CNV12
105
4
Foreword
The CIFOR Permanent Sample Plots (PSP) were established in 1998
in conjunction with the Phase 1 of the ITTO Project dealing with
sustainable forest management, integrated land use, reduced-impact
logging, biodiversity in the production forest, and cultural and social
aspects of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes, long term
growth and yield and to provide critical data for evaluation of ecological
model. For silvicultural purposes PSP supply data on, diameter and
volume increment as well as stand structure dynamics.
I therefore appreciate those who have given strong initiative and
support in the establishment of PSP in Malinau Research Forest and
continuously measure those plots. My utmost appreciation goes also
to our partners: PT INHUTANI II for logging activities, BIOMA, the
Malinau District Office for administrative research arrangements and
the Indonesian Institute of Sciences (LIPI) for scientific identification
of trees I also wish to express my sincere thanks to Dr. Kuswata
Kartawinata, Dr. Plinio Sist of Cirad Foret, Dr. Kade Sidiyasa of
FORDA and Hari Priyadi who maintain their continuing interest by
revisiting the plots and confirming the accuracy of the identity of trees
found in the plots.
Dr Kartawinata and I take the initiative to publish the guidebook to
satisfy the growing interests and needs of visitors visiting our Seturan
camp. More and more people think highly of the importance of these
plots to understand the secret of forest growth and the diversity of
trees occurring inside the permanent plots.
This initiative is not possible without continuous support of ITTO in
providing fund for both research and publication of this guide. It is
my sincerest hope that this little guide will benefit any one who is
concerned with the future of the tropical forest.
Bogor, May 2006
Dr. Petrus Gunarso
Malinau Research Forest Coordinator
5
Introduction
A set of Permanent Sample Plots (PSP) was established in the lowland
mixed dipetrocarp forest within PT Inhutani II concession at Malinau
(2o45’ – 3o15’N, 116o30’), East Kalimantan (Figure 1). They were
designed for comparative studies of conventional logging (CNV) and
reduced-impact logging (RIL) techniques under the ITTO Project PD
12/97 Rev.1 (F), Forest, Science and Sustainability. Bulungan Model
Forest. The Malinau concession is located about 30 km to the East
of CIFOR’s Bulungan Research Forest in a forest area with elevation
between 100 m to 300 m above sea level. The terrain is undulating
with slopes ranging from 10% to 70%. The mean annual rainfall in the
area is 3.790 mm, with the number of wet months of more than 9
months per year.
This pocket guideline is intended to give visitors to PSPs information
on how to access to the location of plots and distribution of trees with
dbh ≥ 20cm in each plot. The CIFOR’s Seturan station is used as a
starting point to travel to the plots.
6
Figure 1. Malinau Research Forest Map
Figure 2. Location of Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 7
8
The Plots
The objective of setting up the permanent sample plots is to monitor
long-term impact of logging on forest dynamic, growth and yield.
The plot locations were selected randomly according to the density
of harvestable timber trees (diameter at breast height or dbh ≥ 60cm)
prior to logging using the tree maps and inventory data in the logging
compartments prepared by CIFOR and PT INHUTANI II. A field
checking was carried out before delimiting each plot to asses the
efective density of harvestable trees. Between June 1998 and September
1998, 12 one-hectare plots were set up in Petak (compartment) 28/29
(Figure 2).
Each one-hectare plot (100 m x 100 m) was horizontally corrected
during its establishment. The plot was divided into 25 subsquares
(20m x 20m), each being divided into 4 elementary squares (a,b,c,d,
in Figure 3 below). At 20 m intervals throughout the plot, PVC piping
was driven into the soil, and a metal tag attached to indicate the sub
square’s coordinates within the plot. A total of 36 PVC stakes were
placed throughout the plot. A wood panel indicating the block and
the number of the plot marks the centre of each. Three control plots
(CNV 04, CNV 08, CNV 12,) were designated and in each one, a 50 m
buffer zone was set up to avoid edge effect caused by logging activities.
After logging, all plots were re-grided (every subsquare 20m x 20 m
materialised by rope) and permanently marked in the field with iron
wood/ulin poles located at each corner.
9
Access to a Plot
Visitors are advised to refer to the first map (Figure 4) to get an
overview on how the plots are spatially distributed. The plots are
grouped based on their route. The routes follow the best walking way
to go to each plot, that is the available skidding road in the field.
To enter a route, visitors should find direction sign in the field posted
on the entrance of each route and follow a skidding road directed by
the sign. Zoomed maps of the routes are given in Figures 4.
Figure 3. Permanent Sample Plot structure (PVC piping is located at
each number)
20m
31 0
20m
20m
c
b
a
c
b
a
26
25 0
c
a
b
c
a
b
d
c
b
a
c
c
d
c
d
c
b
a
b
a
b
a
b
d
c
d
a
b
d
b
d
c
b
a
West
East
South
17
17
18
c
a
b
a
b
d
c
d
c
d
c
b
a
b
a
b
a
9
10
9
11
3
3 0
11
10
2
North
16
d
8
2 0
24
12
4
4 0
5
5 0
20m
a
1 0
23
16
c
1
23
22
15
8
a
d
d
d
7 0
b
b
20m
b
a
30
22
15
7
d
a
14
a
c
b
21
14
c
d
29
a
21
d
b
29
c
20
c
a
d
20
13
b
28
d
20m
c
13 0
a
c
28
d
19
a
b
36
d
20m
c
19 0
c
27
d
35
d
27
26
20m
34
d
20m
d
25
a
20m
33
32
c
b
6 0
10
Plant Identification
Trees with dbh ≥ 20 cm and dbh ≥ 10 cm were identified in the field by
botanists of the Herbarium Bogoriense, Puslitbang Biologi-LIPI, Bogor.
Voucher specimens were collected for further identification and are
stored at the Herbarium Bogorriense. The identification is not final,
hence any corrections are welcome and will be very much appreciated.
The nomenclature follows T.C. Whitmore, I.G.M. Tantra and U. Sutisna
(Editor), Tree Flora of Indonesia: Check List for Kalimantan, Agency
for Forestry Research and Development, Ministry of Forestry, Bogor,
and C.G. G.J. Van Steenis, W.J.O. De Wilde, C. Kalkman, etc. (Editor),
Flora Malesiana Serie I, Volumes 1-15, 1948-2001.
CIFOR RESEARCH STATION
TPN 1
FIRE TOWER
BRIDGE
PSP
RIVER
ROAD
Legend
an
tur
Se
CNV07
CNV03
CNV02
CNV08
CNV04
CNV01
CNV12
CNV10
CNV09
CNV05
CNV06, CNV01, CNV04, CNV07
CNV05, CNV11, CNV09, CNV12, CNV03
CNV15, CNV14, CNV02, CNV10, CNV08, CNV13, CNV16
CNV06
CNV11
Route #1
Route# 2
Route# 3
Figure 4. Distribution of Permanent Sample Plots in Petak (Compartment) # 28 - 29 (Conventional logging treatment)
A Field Guide to the Permanent Sample Plots • 11
100
80
60
40
20
0
0
0
2
1
79
82
83
77
4
76
78
80
9
10
75
74
5
7
88
90
153
163
20
20
12
14
13
69
68
93
15
67
40
64
40
17
19
97 98
100
22 21
65
66
94
95
96
169
141
143 142
168
144
148 145
146
147
167
149
150
165
151
164
91
92
11
70
72 71
87
162
86
158
161
73
89
8
85
157
160
81
84
159
18
24
25
23
63
99
27
60
28
101
26 60
61
176
59
110
109
33
31
56 58
54
111
30
34
136
50
119
35
36
49
80
114
112
44
43
122
42
46
48
127
128
129
191
39
41
47
123
121
125
124
120
132
133
131
130
188
187 189
190
185 186
45
184
183
135
113
80
182
117 116
180
57 55
52 51
53
179
181
108
178
177
107
103
102
105 106
104
139
173
174 175
60
38
40
100
193
192
194
100
100
80
60
40
20
0
Figure 5. Distribution of Trees in Plot CNV01
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Apocynaceae
Annonaceae
Verbenaceae
Ulmaceae
Theaceae
Sterculiaceae
Simarubaceae
Sapotaceae
Sapindaceae
Rosaceae
Podocarpaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lauraceae
FAMILY
Anacardiaceae
Icacinaceae
CNV01
12 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 13
Table 1. List of Trees in Plot CNV01
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea parvifolia Dyer
Shorea fallax Meijer
Unidentified
Diospyros sp. 1
Parashorea smythiesii Wyat-Smith ex Ashton
Unidentified
Shorea parvifolia Dyer
Palaquium rostratum (Miq.) Burk.
Shorea pauciflora King
Dipterocarpus stellatus Vesque
Shorea parvifolia Dyer
Dacryodes rostrata H.J.Lam forma pubescens
Shorea beccariana Burk.
Lithocarpus gracilis (Korth.) Soepadmo
Syzygium sp.
Dead
Alseodaphne paludosa Gamble
Gluta wallichii (Hk.f.) Ding Hou
Canarium caudatum King
Myristica beccarii Warb
Licania splendens (Korth.) Prance
Shorea sp.
Syzygium sp.
Licania splendens (Korth.) Prance
Sindora sp.
Melanochyla auriculata Hook.f.
Shorea fallax Meijer
Parashorea smythiesii Wyat-Smith ex Ashton
Dead
Diospyros sp.
Neesia cf. altissima Bl.
Lithocarpus nieuwenhuisii (V.Seem) A. Camus
Alseodaphne havilandii (Gamble) Kosterm.
Shorea fallax Meijer
Pternandra rostrata (Cogn.) Ohwi
Lithocarpus blumeanus (Korth.) Rehd.
Dead
14 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Livistona sp.
Livistona sp.
Tetramerista glabra Miq.
Madhuca sp.
Tetramerista glabra Miq.
Pternandra caerulescens Jack.
Lithocarpus benetii (Miq.) Rehd.
Dacryodes rugosa (Bl.) H.J. Lam
Diospyros borneensis Hiern)
Lithocarpus urceolaris (Jack) Merr.
Xylopia altissima Boerl.
Syzygium tawahense (Korth) Merr. & Perry
Shorea parvifolia Dyer ssp.parvifolia
Alseodaphne havilandii (Gamble) Kosterm.
Palaquium rostratum (Miq.) Burk.
Polyalthia sumatrana King
Aglaia cordata Hiern.
Livistona sp.
Parashorea smyethiesi Wyatt-Smith exAshton
Cleistanthes myrianthus Krz.
Vatica rassak (Korth.) Blume
Aporusa falcifera Hk.f.
Shorea ovalis (Korth.) Bl.
Hydnocarpus cf. kunstleri (King) Warb.
Shorea ovalis (Korth.) Bl.
Diospyros ellipticifolia Merr.
Syzygium perpuncticulatum (Merr.) Merr. & Perry
Shorea sp.
Litsea elliptica Bl.
Myristica maxima Warb.
Bhesa paniculata Arn.
Ryparosa acuminata Merr.
Sarcotheca diversifolia (Miq.) Hall.f.
Parashorea smythiesii Wyat-Smith ex Ashton
Heritiera sp.
Artocarpus lanceifolius Roxb.
Garcinia nervosa Miq.
A Field Guide to the Permanent Sample Plots • 15
Table 1. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Syzygium sp.
Actinodaphne glabra Bl.
Stemonurus grandifolius Becc.
Syzygium endertii Merr. & Perry
Artocarpus lanceifolius Roxb.
Diospyros
Irvingia malayana Oliver
Castanopsis philippinensis Vidal
Dillenia exelsa (Jacq.) Gilg.
Parashorea smythiesii Wyat-Smith ex Ashton
Lithocarpus ewyckii (Korth) Rehd.
Nephelium maingayi Hiern.
Dillenia exelsa (Jacq.) Gilg.
Diospyros sp.
Parashorea smythiesii Wyat-Smith ex Ashton
Diospyros sp.
Dehaasia cf. firma Bl.
Podocarpus becarii Parl.
Canarium denticulatum Bl. ssp. denticulatum
Cephalomappa lepidotula Airy Shaw
Tetramerista glabra Miq.
Garcinia maingayi Hook.f.
Popowia pisocarpa (Bl.) Endl.
Castanopsis motleyana King
Shorea ovalis Korth.) Bl.
Cleistanthus myrianthus Krz.
Koompassia malaccensis Maing. ex Benth.
Teijsmaniodendron simplicifolium Merr.
Diospyros macrophylla Bl.
Dyera costulata (Miq.) Hk.f.
Sindora leiocarpa Backer
Teijsmaniodendron simplicifolium Merr.
Syzygium sp.
Shorea agami Ashton
Dipterocarpus humeratus Sloot.
Heritiera simplicifolia (Mart.) Kosterm.
Syzygium tawahense (Korth.) Merr. & Perry
16 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Coccoceras borneense J.J. Smith
Teijsmaniodendron simplicifolium Merr.
Sarcotheca diversifolia (Miq.) Hall.f.
Teijsmaniodendron simplicifolium Merr.
Teijsmaniodendron simplicifolium Merr.
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Dillenia eximia Miq.
Girroniera nervosa Planch.
Dyera costulata (Miq.) Hk.f.
Dacryodes sp.
Scaphium macropodum (Miq.) Beumee ex Heyne
Alseodaphne insignis Gamble
Teijsmaniodendron simplicifolium Merr.
Heritiera simplicifolia (Mart.) Kosterm.
Dead
Palaquium ferox H. J. Lam
Dacryodes sp.
Palaquium calophyllum Pierre ex Burck
Heritiera simplicifolia (Mart.) Kosterm.
Shorea parvifolia Dyer
Teijsmaniodendron simplicifolium Merr.
Tetramerista glabra Miq.
Dead
Palaquium gutta (Hook.f.) Bail
Syzygium chlorantum (Dut.) Merr. & Perry
Baccaurea sumatrana M.A.
Herritiera javanica (Bl.) Kosterm
Cleistanthus myrranthus (Krz.
Dead
Palaquium ferox H.J. Lam
Scaphium macropodum (Miq.) Beumée ex Heyne
Knema conferta (King) Warb
Artocarpus odoratisima Blanco
Oncosperma horridum Scheff
Oncosperma horridum Scheff
Scaphium macropodum (Miq.) Beumée ex Heyne
Canarium denticulatum Bl. ssp. denticulatum
A Field Guide to the Permanent Sample Plots • 17
Table 1. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Santiria oblongifolia Bl.
Syzygium sp.
Shorea ovalis (Korth.) Bl.
Castanopsis hypophoenicea (V.Sleum.) Supadmo
Bhesa paniculata Arn.
Shorea ovalis (Korth.) Bl.
Macaranga pruinosa (Miq.) Muell. Arg
Dead
Artocarpus kemando Miq
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea parvifolia Dyer
Shorea ovalis (Korth.) Bl.
Baccaurea sumatrana M.A.
Durio sp.
Artocarpus elasticus Reinw.
Licania splendens (Korth.) Prance
Heritiera simplicifolia (Mart.) Kosterm.
Alseodaphne insignis Bamble
Vatica vinosa Ashton
Polyalthia rumphii Merr.
Vatica vinosa Ashton
Dipterocarpus stellatus Vesque
Sindora leiocarpa Backer
Artocarpus kemando Miq.
Durio oxleyanus Griff.
Heritiera simplicifolia (Mart.) Kosterm.
Licania splendens (Korth.) Prance
Myristica beccarii Warb.
Aglaia cordifolia Merr.
Nephelium uncinatum Leenh.
Shorea pinanga Scheff.
Dialium sp.
Dyera costulata (Miq.) Hk.f.
Shorea pinanga Scheff.
Knema cinerea (Poir.) Warb.
Knema cinerea (Poir.) Warb.
Licania splendens (Korth.) Prance
18 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
186
187
188
189
190
191
192
193
194
Species
Teijsmaniodendron simplicifolium Merr.
Dehaasia firma Bl.
Canarium denticulatum Bl. ssp. denticulatum
Shorea macroptera Dyer
Alseodaphne sp.
Syzygium tawahense (Korth.) Merr. & Perry
Garcinia sp.
Shorea fallax Meijer
Syzygium tawahense (Korth.) Merr. & Perry
100
80
60
40
20
0
0
0
239
1
14
112
9
253
20
105
107
136
223
224
222
258
40
221
261
217
267
214
60
207
200
129
17
16
18
101
132
40
99
100
33
96
21
31
22
30
34
98
140
23
29
24
28
35
91
60
27
36
90
25
92
93
38
155
154 89
156
37
47
275
48
46
86
85
44
45
78
80
79
81
82
164
80
280
49
67
66
182
64
75
169
168
173
52
53
62
71
56
100
55
57
69 68
59
70
171
72
74
170
172
174
175
176
177
180
183
178
60
63 61
50
51
77
76
166
191
190
189
286
100
283
285
284
281
282
184
185
188
187 186
278
165
277
80
41 42
39
40
84
83
192
204
199
193
274
88
194
273
203
195
198
209
197
270
269
271 272
212
210
213
211
264
265
266
218
220
263
262
260
215
208
216 145 147
148
138 134 135
149
144
139
146
150 161 159
143 142
128 133
141
152
225
257
256
259
254
255
104
130
131
108
126
229
227 228
250
230
119
7
20
252
251
121
124
231
232
249
246
233
24 5
6
3
117
113 12
13 11 10
114
116
115
122
125
236
248
245
244
243
234
247
235
240
242
100
80
60
40
20
0
Figure 6. Distribution of Trees in Plot CNV02
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Crypteroniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Tiliaceae
Thymelaeaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Sabiaceae
Rubiaceae
Rosaceae
Proteaceae
Polygalaceae
Oxalidaceae
Olacaceae
Ochnaceae
Myrtaceae
Myristicaceae
Meliaceae
Lauraceae
FAMILY
Anacardiaceae
Flacourtiaceae
CNV02
A Field Guide to the Permanent Sample Plots • 19
20 • A Field Guide to the Permanent Sample Plots
Table 2. List of Trees in Plot CNV02
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Anisoptera sp.
Castanopsis oviformis Soepadmo
Castanopsis oviformis Soepadmo
Diospyros sp.
Baccaurea minor Hook.f.
Litsea grandis (Wall. ex Nees) Hook.f.
Baccaurea minor Hook.f.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pimeleodendron papaveroides J.J.S.
Shorea parvifolia Dyer
Dacryodes rostrata H.J.Lam
Dacryodes rostrata H.J.Lam
Sterculia macrophylla Vent.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Koordersiodendron pinnatum (Blanco) Merr.
Saraca hullettii Prain.
Lithocarpus urceolaris (Jack.) Merr.
Knema latifolia Warb.
Shorea pauciflora King
Nephelium uncinatum Leenh.
Knema cinerea (Poir.) Warb.
Dysoxylum sp.
Santiria oblongifolia Bl.
Alseodaphne ceratoxylon Kosterm.
Sarcotheca diversifolia (Miq.) Hall.f.
Dehaasia firma Bl.
Irvingia malayana Oliv.
Melanochyla beccariana Oliv.
Dead
Castanopsis tungurut (Bl.) A.DC.
Sindora sp.
Alseodaphne ceratoxylon Kosterm.
Parashorea smythiesii Wyatt-Smith ex Ashton
Parashorea smythiesii Wyatt-Smith ex Ashton
A Field Guide to the Permanent Sample Plots • 21
Table 2. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Santiria laevigata Bl.
Vatica rassak (Korth.) Blume
Alseodaphne cuneata (Bl.) Kosterm.
Myristica sp.
Polyalthia sp.
Dead
Syzygium sp.
Lithocarpus ulceolaris (Jack) Merr.
Canarium caudatum King
Shorea macroptera Dyer.
Xanthophyllum parvum Chod.
Timonius ovalis Boerl
Shorea parvifolia Dyer
Maccaranga gigantea M.A.
Shorea parvifolia Dyer
Dehaasia firma Bl.
Dead
Syzygium sp.
Polyalthia glauca Boerl.
Shorea seminis Slooten
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Shorea macroptera Dyer
Artocarpus odoratissimus Blanco
Knema intermedia Warb.
Helicia fuscotomentosa Suesseng
Dead
Polyalthia lateriflora (Bl.) King
Shorea exelliptica Meijer
Aporusa antennifera (Airy Shaw) Airy Shaw
Santiria laevigata Bl. f. laevigata
Lithocarpus bennetii (Miq.) Rehd.
Dyera costulata (Miq.) Hk.f.
Agathis borneensis Warb.
Dead
Macaranga pruinosa M.A.
22 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Ochanostachys amentacea Mast.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Elaeocarpus grifithii A. Gray
Syzygium sp.
Vatica umbonta (Hook.f.) Burk.
Dyera costulata (Miq.) Hk.f.
Diospyros kunstleri King & Gamble
Shorea parvifolia Dyer.
Myristica beccarii Warb.
Baccaurea deflexa M.A.
Pometia pinnata Forst.
Dead
Melanochyla beccariana Oliv.
Chisocheton sp.
Syzygium sp.
Shorea parvifolia Dyer
Shorea pauciflora King
Shorea parvifolia Dyer
Dead
Shorea parvifolia Dyer
Shorea pauciflora King
Unidentified
Koompassia malaccense Maing.. ex Benth.
Aquilaria malaccensis Lam.
Koompassia excelsa Taub.
Knema glauca (Bl.) Warb.
Dead
Dead
Pentace erectinervia Kosterm.
Dillenia excelsa (Jack.) Gilg.
Albizia sp.
Knema conferta (King) Warb.
Macaranga hypoleuca (Richb. f & Zoll.) M.A.
Uidentified
Dead
Saraca hulletii Prain
A Field Guide to the Permanent Sample Plots • 23
Table 2. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Syzygium operculata Gamb.
Eugeissona utilis Becc.
Mangifera swintonioides Kosterm.
Saraca hulletii Prain
Shorea parvifolia Dyer
Mangifera swintonioides Kosterm.
Knema cinerea (Poir.) Warb
Livistona sp.
Saraca hulletii Prain
Macaranga hypoleuca (Richb. f & Zoll.) M.A.
Shorea parvifolia Dyer
Unidentified
Shorea parvifolia Dyer
Diospyros ellipticifolia Merr.
Syzygium sp.
Dead
Cococeras borneense J.J. Smith
Lithocarpus nieuwenhuisii (v. Seem) A. Camus
Knema laurina (Bl.) Warb.
Baccaurea kunstlerii King
Shorea parvifolia Dyer
Koompassia malaccensis Maing.. ex Benth.
Shorea parvifolia Dyer
Xanthophyllum amoenum Chod.
Baccaurea angulata Merr.
Unidentified
Canarium denticulatum Bl. ssp. denticulatum
Shorea parvifolia Dyer ssp. velutina
Knema cinerea (Poir.) Warb.
Ptychophyxys bacciformis Croiz.
Ptychophyxys bacciformis Croiz.
Koompassia malaccensis (Clarke) Breh.
Macaranga hypoleuca (Reichb.f. & Zoll.) M.A.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pouteria malaccensis (Clarke) Breh.
Knema sp.
24 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Lythocarpus ewyckii (Korth.) Rehd.
Cephalomappa lepidotula Airy Shaw
Shorea parvifolia Dyer
Koompassia malaccensis Maing.. ex Benth.
Girroniera nervosa Planch.
Shorea pauciflora King
Artocarpus kemando Miq.
Artocarpus lanceifolius Roxb.
Unidentified
Unidentified
Lythocarpus ewyckii (Korth.) Rehd.
Macaranga hypoleuca (Reichb.f. & Zoll.) M.A.
Artocarpus lanceifolius Roxb.
Dead
Mangifera swintonoides Kosterm.
Crypteronia griffithii Clarke
Payena endertii H.J.Lam
Glochidion macrocarpum Bl.
Dead
Polyalthia sclerophylla Hook.f. ex Th.
Dialium trifoliatum de Wit
Garcinia dioica Bl.
Mangifera swintonioides Kosterm.
Palaquium calophyllum Pierre ex Burck
Lithocarpus ewyckii (Korth.) Rehd.
Shorea parvifolia Dyer
Dacryodes rostrata H.J. Lam
Dacryodes rostrata H.J. Lam
Shorea pauciflora King
Durio oxleyanus Griff.
Dead
Shorea exelliptica Meijer
Knema latericia Elmer
Shorea sp.
Buchanania sp.
Shorea exelliptica Meijer
Scaphium macropodum (Miq.) Buemee ex Heyne
A Field Guide to the Permanent Sample Plots • 25
Table 2. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Knema sp.
Dialium trifoliatum De Wit
Atuna excelsa (Jack.) Kosterm.
Dialium trifoliatum De Wit
Koordersiodendron pinnatum (Blanco) Merr.
Dacryodes rostrata H.J. Lam
Drypetes kikir Airy Shaw
Shorea pauciflora King
Mangifera swintonioides Kosterm.
Helicia serrata (R.Br.) Bl.
Dead
Baccaurea edulis Merr.
Knema latericia Elmer.
Durio lanceolatus Mast.
Dialium platycephalum Backer
Lithocarpus ewyckii (Korth.) Rehd.
Mangifera swintonioides Kosterm.
Sindora leiocarpa Backer ex Heyne
Oncosperma horridum Scheff.
Canarium denticulatum Bl. ssp. denticulatum
Dipterocarpus sp.
Crypteronia griffithi Clarke
Hydnocarpus anomala (Merr.) Sleumer
Dialium platycephalum Backer
Artocarpus kemando Miq.
Knema furfuracea (Hk.f. ex Th.) Warb.
Cratoxylum formosum Benth. & Hook. f. ex Dyer
Vatica umbonata (Hook.f.) Burk.
Dipterocarpus
Artocarpus lanceifolius Roxb.
Palaquium calophyllum Pierre ex Burck
Shorea parvifolia Dyer
Parashorea amythiesii Wyatt-Smith ex Ashton
Dead
Pouteria malaccensis (Clarke) Breh
Diospyros sp.
Meliosma nitida
26 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Syzygium sp.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Unidentified
Mangifera magnifica Kochumen
Syzygium sp.
Nephelium eriobetalum Miq.
Polyalthia lateriflora (Bl.) King
Bhesa paniculata Arm.
Garcinia dioica Bl.
Vatica micrantha Slooten
Dacryodes incurvata (Engl.) H.J.Lam.
Pometia pinnata Forst.
Dacryodes costata (Benn.) H.J. Lam
Artocarpus integer (Thunb.) Merr.
Vatica micrantha Slooten
Cococeras borneensis M.A.
Dipterocarpus cornutus Dyer
Dead
Dipterocarpus confertus Sloot.
Agathis borneensis Warb.
Cococeras borneensis M.A.
Baccaurea sp.
Beilschmiedia sp.
Diopyros ellipticifolia Merr.
Agathis borneensis Warb.
Diospyros everetii Merr.
Bhesa paniculata Arn.
Elateriospermum tapos Bl
Ochanostachys amentacea Mast.
Cleistanthus myrianthus (Hassk.) Kurz.
Palaquium sp.
Sarcotheca diversifolia (Miq.) Hall.f.
Palaquium ottolanderi K. ex V.
Prunus sp.
Palaquium calophyllum Pierre ex Burck
Dysoxylum sp.
A Field Guide to the Permanent Sample Plots • 27
Table 2. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
Species
Cephalomappa beccariana Baill.
Sindora leiocarpa Backer
Dipterocarpus humeratus Sloot.
Elateriospermum tapos Bl.
Madhuca cf. rupicola H.J.Lam
Dacryodes rugosa H.J.Lam var. rugosa
Xanthophyllum vittelinum (Bl.) Dierr.
Xylopia malayana Hook. f. & Thoms.
Dead
Plectronia glabra Benth. & Hook. ex Kurz
Shorea macroptera Dyer.
Brownlowia sp.
Syzygium sp.
Dacryoides rostrata H.J. Lam
Shorea sp.
Dipterocapus stellatus Vesque
Unidentified
Shorea macroptera Dyer.
Nauclea subdita Steud.
Dead
Shorea parvifolia Dyer.
Dipterocapus stellatus Vesque
Shorea parvifolia Dyer.
Helicia serrata Bl.
Dipterocapus stellatus Vesque
Gymnacranthera paniculata Warb.
Gymnacranthera paniculata Warb.
100
80
60
40
20
0
0
0
1
3
94
180
185
178
186
7
6
95
93
91
176
184
5
90
98
8
103
20
4
12
11
17
86
85
81
14 13
83
82
105
84
87
15
10
9
89
193
77
40
76
78
80
107
74
116
75
21
24
26
70
28
68
111
163
162
69
22
23
19
20
25
71
200
199
110
115
198
73 72
108
40
106
172
171
166
165
197
195
194
196
18
79
174
173
169
168 167
190
191
170
192
189
102 101
88
99
100
181
182
183
20
112
113
62
63
65
34
123
151
59
58
57
126
124
122
152
61
120
66 64
118
30
60
121
153
154
206
80
32
80
36
37
54
39
40
42
53
141
217
100
100
48
47
46
45
44
49
133
135
137
139 138
142
140
215
132
50
52
51
128
130
55
129
131
149
147
146
144 143
212
148
38
60
56
127
150
145
159
158 160
205
207
209 211
210
208
204
156 157
202
117
29
114
161
203
201
60
100
80
60
40
20
0
Figure 7. Distribution of Trees in Plot CNV03
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Annonaceae
Ulmaceae
Tiliaceae
Thymelaeaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Proteaceae
Oleaceae
Olacaceae
Myrtaceae
Myristicaceae
Meliaceae
FAMILY
Anacardiaceae
Lauraceae
CNV03
28 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 29
Table 3. List of Trees in Plot CNV03
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Polyalthia glauca Boerl.
Dead
Croton laevifolim Bl.
Artocarpus kemando Miq.
Eugeissona utilis Becc.
Shorea ovalis (Korth.) Bl.
Myristica maxima Warb.
Coccoceras borneense J.J. Smith
Beilschmiedia bankae Kosterm.
Syzygium splendens (Bl.) Merr. & Perry
Gonystylus brunnescens Airy Shaw
Baccaurea
Vatica rassak (Korth.) Blume
Syzygium splendens (Bl.) Merr. & Perry
Vatica oblongifolia Hook.f.
Unidentified
Vatica rassak (Korth.) Blume
Syzygium splendens (Bl.) Merr. & Perry
Parashorea smythiesii Wyatt-Smith ex Ashton
Cephalomappa lepidotula Airy Shaw
Cephalomappa lepidotula Airy Shaw
Vatica micrantha Slooten
Coccoceras borneense J.J. Smith
Shorea ovalis (Korth.) Bl.
Dillenia excelsa (Jacq.) Gilg.
Shorea parvifolia Dyer
Dead
Cephalomappa lepidotula Airy Shaw
Dyospiros sp.
Syzygium operculata Gamble
Dead
Parashorea sp.
Dead
Vatica rassak (Korth.) Blume
Turpinia sphaerocarpa Hassk.
Litsea noronhae Bl.
Coccoceras borneense J.J. Smith
30 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Shorea pauciflora King
Durio kuteyensis Becc.
Dysoxylum sp.
Pternandra rostrata (Cogn.) Ohwi
Artocarpus lanceifolius Roxb.
Dead
Dysoxylum hexandrum Merr.
Scaphium macropodum (Miq.) Beumée ex Heyne
Lithocarpus ewyckii Korth.) Rehd.
Lithocarpus ewyckii Korth.) Rehd.
Myristica malaccensis Hk.f.
Aglaia cordata Hiern.
Lithocarpus ewyckii (Korth.) Rehd.
Shorea ovalis Korth. Bl.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Gymnacranthera paniculata Warb.
Pentace laxiflora Merr.
Bhesa paniculata Arn.
Artocarpus anysophyllus Miq.
Chionanthes cuspidatus Bl.
Coccoceras borneense J.J. Smith
Chionanthus sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Baccaurea sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Drypetes longifolia Pax. & Hoffm.
Shorea ovalis (Korth.) Bl.
Syzygium sp.
Dead
Coccoceras borneense J.J. Smith
Baccaurea sp.
Dipterocarpus stellatus Vesque
Diospyros malabarica (Desr.) Kostel.
Pentace discolor Merr.
Herritiera javanica (Bl.) Kosterm.
Coccoceras borneense J.J. Smith
A Field Guide to the Permanent Sample Plots • 31
Table 3. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Coccoceras borneense J.J. Smith
Coccoceras borneense J.J. Smith
Shorea maxwelliana King
Licania spendens (Korth.) Prance
Chionanthus cuspidatus Bl.
Parashorea smythiesii Wyatt-Smith ex Ashton
Coccoceras borneense J.J. Smith
Shorea parvifolia Dyer
Lithocarpus nieuwenhuisii (V.Seem) A.Camus
Shorea parvifolia Dyer
Syzygium sp.
Knema cinerea (Poir) Warb.
Licania splendens (Korth.) Prance
Vatica rassak (Korth.) Blume
Castanopsis hypophoenica (V.Seem) Soepadmo
Diospyros malabarica (Desr.) Kostel.
Vatica rassak (Korth.) Blume
Planchonia valida (Bl.) Bl.
Vatica micrantha Slooten
Shorea fallax Meijer
Shorea pauciflora King.
Syzygium splendens (Bl.) Merr. & Perry
Shorea fallax Meijer
Licuala sp.
Licuala sp.
Neonauclea purpurascens Moore
Shorea ovalis (Kort.) Bl.
Knema cinerea (Poir.) Warb.
Cratoxylum sumatranum (Jack.) Bl.
Dead
Aporusa falcifera Hk.f.
Shorea pauciflora King
Castanopsis javanica (Bl.) A.DC.
Dipterocarpus stellatus Vesque
Dead
Palaquium sumatranum Burck.
Syzygium sp.
32 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Drypetes longifolia Pax. & Hoffm.
Durio carrinatus Mast.
Syzygium sp.
Dysoxylum hexandrum Merr.
Dehaasia firma Bl.
Parashorea smythiesii Wyatt-Smith ex Ashton
Ochanostachys amentacea Mast.
Dead
Dacryodes rogosa H.J.Lam
Coccoceras borneense J.J. Smith
Knema furfuracea (Hk. f. ex Th.) Warb.
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea pinanga Scheff.
Dead
Shorea beccarina Burk.
Shorea parvifolia Dyer
Knema intermedia (Bl.) Warb
Shorea ovalis (Kort.) Bl.
Shorea pauciflora King
Artocarpus kemando Miq.
Koompassia malaccensis Maing. ex Benth
Dialium patens Backer
Dead
Palaquium rostratum (Miq.) Burk.
Dead
Shorea pauciflora King
Parashorea smythiesii Wyatt-Smith ex Ashton
Syzygium splendens (Bl.) Merr. & Perry
Lithocarpus blumeanus (Korth.) Rehd.
Gymnacranthera paniculata Warb.
Knema furfuracea (Hk.f. ex Sinel.) Warb
Knema laurina (Bl.) Warb.
Vatica rassak (Korth.) Blume
Durio oxleyanus Griff.
Knema laurina (Bl.) Warb.
Pimeleodendron papaveroides J.J.S.
Parashorea smythiesii Wyatt-Smith ex Ashton
A Field Guide to the Permanent Sample Plots • 33
Table 3. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea pinanga Scheff.
Syzygium angustifolius (Miq.) Hats.
Drypetes sp.
Pometia pinnata Forst.
Durio testudinarum Becc.
Parashorea smythiesii Wyatt-Smith ex Ashton
Dead
Diospyros sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Knema glauca (Bl.) Warb.
Garcinia parvifolia Miq.
Knema glauca (Bl.) Warb.
Lithocarpus conocarpus (Oudem) Rehd
Shorea seminis (de Vriese) Slooten
Shorea pinanga Scheff
Dead
Coccoceras borneense J.J. Smith
Shorea rubra Ashton
Coccoceras borneense J.J. Smith
Vatica micrantha Slooten
Baccaurea sp.
Cleisthanthus myrianthus Krz.
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea rubra Ashton
Pterocymbium tubulatum (Mast.) Pierre
Alseodaphne ceratoxylon Kosterm.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer
Actinodaphne glabra Bl.
Pterocymbium tubulatum (Mast.) Pierre
Dead
Livistona
Shorea parvifolia Dyer
Payena accuminata (Bl.) Pierre
Palaquium rostratum (Miq.) Burk.
Gymnacranthera paniculata Warb.
Diospyros everetii Merr.
34 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
Species
Knema furfuracea (Hk.f.ex. Th.) Warb.
Dead
Livistona sp.
Dialium modustum (V. Stenis) Stegaert
Calophyllum soulatri Burm.
Cratoxylum formosum (Jack) Dyer
Syzygium splendens (Bl.) Merr. & Perry
Aglaia cf. pseudolansium Kosterm.
Shorea pauciflora King
Mangifera quadrifida Jack.
Dialium modestum (V. Steenis) Stegaert.
Pentace discolor Merr.
Syzygium splendens (Bl.) Merr. & Perry
Aporusa falcifera Hk.f.
Cleisthanthus myrianthus Krz.
Castanopsis javanica (Bl.) A.DC.
Celtis nigrecens (Miq.) Planch.
Sindora leiocarpa Backer
Diospyros cf. oblonga Wall. ex. G. Don
Prunus sp.
Coccoceras borneense J.J. Smith
Shorea fallax Meijer
Castanopsis javanica (Bl.) A.DC.
Helicia petiolaris Benn.
Coccoceras borneense J.J. Smith
Myristica sp.
Diospyros ellipticifolia Merr.
Aporusa falcifera Hk.f.
Gardenia tubifera Wall.
Diospyros oblonga Wall. ex. G. Don
Calophyllum soulatri Burm.
Diospyros oblonga Wall. ex. G. Don
100
80
60
40
20
0
0
0
2
1
5
84
83
95
94
162
90
3
6
82
81
86
88
87
89
79
91 96
92
7 8
164
163
169
171
174
168
161
165
166
167
172
20
159
20
156
76
9
10
11
100
99
101 102
77
98
160
157
180
182
179
158
78
170
178
177
176
175
13
12
75
103
155
153
183
40
192
190
14
16
15
21
68
17
40
73
74
71
70
69
105 108
107
106
151
191
196
18
67
145
144
28
25
63
26
23
27
60
62
66
19
20
22
61
64
119
116
197
60
60
29
59
58
120
30
57
52
51
203
205
34
31
32 33
56
53
50
49
125
124
137
134
38
35
80
36 37
54
47
48
136
46
215
44
132
212
220
223
214
45
133
213
218
217
216
209
208
80
138
210
211
207
140 139
201
202
204
206
123
122
121
117
143
147
146
141
200
199
198
115
114 118
109
111
112
110
113
148
149
150
195
186
193
194
104
152
184
187
189
154
185
188
41
43
39
42
100
128
129
130
131
221
222
219
100
100
80
60
40
20
0
Figure 8. Distribution of Trees in Plot CNV04
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Apocynaceae
Annonaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Rhizophoraceae
Proteaceae
Polygalaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lauraceae
FAMILY
Anacardiaceae
Flacourtiaceae
CNV04
A Field Guide to the Permanent Sample Plots • 35
36 • A Field Guide to the Permanent Sample Plots
Table 4. List of Trees on Plot# CNV04
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Aporusa falcifera Hk.f.
Pternandra rostrata (Cogn.) Ohwi
Alstonia angustiloba Miq.
Dead
Memecylon myrsinoides Blume
Syzygium bankense (Hassk.) Merr. & Perry
Dialium trifoliatum de Wit.
Artocarpus kemando Miq.
Shorea pinanga Scheff.
Aglaia cf. gamopetala Merr.
Litsea forstenii (Bl.) Boerl.
Palaquium rostratum (Miq.) Burck.
Dacryodes incurvata (Engl.) H.J.Lam
Sarcotheca diversifolia (Miq.) Hall.f.
Koompassia malaccensis Maing. ex Benth
Syzygium tawahense (Korth.) Merr. & Perry
Knema cinerea (Poir.) Warb.
Baccaurea kunstleri King.
Pometia pinnata Forst.
Dyera costulata (Miq.) Hk.f.
Sindora leiocarpa Backer
Parastemon urophyllus (A.DC.) A.DC.
Ternstroemia aneura Miq.
Dead
Calophyllum lowii Hook.f.
Cephallomappa lepidotula Airy Shaw
Myristica beccarii Warb.
Shorea parvifolia Warb.
Baccaurea bracteata M.A.
Myristica beccarii Warb.
Ternstroemia aneura Miq.
Tetramerista glabra Miq.
Dipterocarpus lowii Hk.f.
Myristica beccarii Warb.
Durio lanceolatus Mast.
Shorea seminis (de Vriese) Slooten
Canarium sp.
A Field Guide to the Permanent Sample Plots • 37
Table 4. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Calophyllum pulcherrimum Wall. ex Planch. & Trian.
Palaquium rostratum (Miq.) Burk.
Dead
Durio carinatus Mast.
Santiria laevigata Bl.f. laevigata
Vatica rassak (Korth.) Blume
Canarium littorale Bl.
Myristica beccarii Warb.
Shorea seminis (de Vriese) Slooten
Parastemon urophyllus (A.DC.) A.DC.
Santiria mollissima Ridl.
Shorea seminis (de Vriese) Slooten
Santiria laevigata Bl.
Syzygium tawahense (Korth.) Merr. & Perry
Xanthophyllum parvum Chod
Dyera costulata (Miq.) Hk.f.
Gymnacranthera contracta Warb.
Dead
Shorea seminis (de Vriese) Slooten
Gymnacranthera contracta Warb.
Horsfieldia sp.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Dehaasia firma Bl.
Shorea ovalis (Korth.) Bl.
Gymnacranthera paniculata Warb.
Gymnacranthera paniculata Warb.
Dead
Knema sp.
Koompassia malaccensis Maing. ex. Benth.
Gluta wallichii (Hk.f.) Ding Hou
Sterculia oblongata R.Br.
Santiria oblongifolia Bl.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea seminis (de Vriese) Slooten
Dialium indum L.
38 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Baccaurea pendula Merr.
Dyera costulata (Miq.) Hk.f.
Syzygium splendens (Bl.) Merr. & Perry
Xylopia caudata Hook. f. & Thoms.
Xylopia caudata Hook. f. & Thoms.
Dead
Santiria laevigata Bl.
Durio sp.
Helicia petiolaris Benn.
Baccaurea sumatrana M.A.
Dead
Gymnacranthera paniculata Warb.
Lophopetalum subobovatum King.
Syzigium fastigiata (Bl.) Merr. & Perry
Cephalomappa lepidotula Airy Shaw
Endiandra ochracea Kosterm.
Baccaurea sp.
Aporusa falcifera Hk.f.
Mastixia trichotoma Bl.
Dehaasia firma Bl.
Aporusa falcifera Hk.f.
Knema cinerea (Poir.) Warb
Dead
Shorea parvifolia Dyer
Parashorea smythiesii Wyatt-Smith ex Ashton
Ixonanthes petiolaris Bl.
Hydnocarpus woodii Merr.
Canarium litorale Bl.
Pternandra rostrata (Cogn.) Ohwi
Memecylon edule Roxb.
Castanopsis motleyana King
Shorea parvifolia Dyer
Syzygium litorale (Bl.) Merr. & Perry
Shorea parvifolia Dyer
Vatica vinosa Ashton
Shorea maxweliana King
Canarium litorale Bl.
A Field Guide to the Permanent Sample Plots • 39
Table 4. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea parvifolia Dyer
Palaquium sp.
Gymnacranthera contracta War.
Nephelium maingayi Hiern.
Shorea agami Ashton
Gymnacranthera contracta War.
Pimeleodendron papaveroides J.J.S.
Dehaasia firma Bl.
Horsfieldia sp.
Tetramerista glabra Miq.
Shorea leprosula Miq.
Shorea agami Ashton
Santiria laevigata Bl.
Xylopia altissima Boerl.
Dead
Madhuca malaccensis (Clarke) H.J.Lam.
Myristica sp.
Syzygium ochneocarpa Merr.
Palaquium rostratum (Miq.) Burk.
Myristica vilosa Warb.
Tetramerista glabra Miq.
Lithocarpus cantleyanus (King ex Hk.f.)Rehd.
Vatica rassak (Korth.) Blume
Dead
Palaquium rostratum (Miq.) Burk.
Diospyros oblonga Wall. ex G. Don
Xylopia altissima Boerl.
Madhuca malaccensis (Clarke) H.J.Lam
Pternandra rostrata (Cogn.) Ohwi
Castanopsis aviformis Soepadmo
Dead
Dacryodes incurvata (Engler) H.J.Lam
Syzygium sp.
Durio carinatus Mast.
Oncosperma horridum Scheff.
Shorea seminis (de Vriese) Slooten
Koordersiodendron pinnatum (Blanco) Merr.
40 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Sarcotheca diversifolia (Miq.) Hall.f.
Tetramerista glabra Miq.
Ternstroemia aneura Miq.
Baccaurea pendula Merr.
Alseodaphne insignis Gamble
Parashorea smythiesii Wyatt-Smith ex Ashton
Aporusa falcifera Hk.f.
Buchanania arborescens Bl.
Madhuca malaccensis (Clarke) H.J.Lam
Endiandra sp.
Nephelium maingayi Hiern.
Pternandra rostrata (Cogn.) Ohwi
Shorea pauciflora King
Pternandra rostrata (Cogn.) Ohwi
Shorea rubra Ashton
Parastemon urophyllus (A.DC.) A.DC.
Lithocarpus pulcher (King.) Markgr.
Dipterocarpus lowii Hk.f.
Sizygium splendens (Bl.) Merr. & Perry
Parishia maingayi Hk.f.
Xylopia malayana Hk.f. ex Th.
Elaeocarpus petiolatus Wall.
Artocarpus anisophyllus Miq.
Shorea parvifolia Dyer
Unidentified
Xylopia malayana Hk.f. ex Th.
Parastemon urophyllus (A.DC.) A.DC.
Diospyros sp.
Castanopsis motleyana King
Palaquium rostratum (Miq.) Burk.
Madhuca malaccensis (Clarke) H.J.Lam.
Sarcotheca diversifolia (Miq.) Hall.f.
Dead
Myristica vilosa Warb.
Jackia ornata Wall.
Dehaasia firma Bl.
Syzigium chloranthum (Duth.) Merr. & Perry
A Field Guide to the Permanent Sample Plots • 41
Table 4. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
223
Species
Semecarpus glaucus Engl.
Pternandra rostrata (Cogn.) Ohwi
Cleisthantus myrianthus Krz.
Castanopsis motleyana King
Baccaurea racemosa (Rinw.) Muel. Arg.
Madhuca malaccensis (Clarke) H.J.Lam.
Cephalomappa lepidotula Airy Shaw
Vatica rassak (Korth.) Blume
Baccaurea edulis Merr.
Baccaurea edulis Merr.
Tetramerista glabra Miq.
Syzigium chloranthum (Duth.) Merr. & Perry
Baccaurea puberula Merr.
Lithocarpus urceolaris (Jack.) Merr.
Ternstroemia aneura Miq.
Baccaurea puberula Merr.
Sterculia foetida L.
Shorea parvifolia Dyer
Shorea seminis (de Vriese) Slooten
Syzygium bankense (Hassk.) Merr. & Perry
Shorea seminis (de Vriese) Slooten
Gymnacranthera paniculata Warb.
Ficus sp.
Syzigium chloranthum (Duth.) Merr. & Perry
Baccaurea sp.
Carallia brachyata (Lour.) Merr.
Xanthophyllum sp.
Tetramerista glabra Miq.
Knema sp.
Baccaurea puberula Merr.
Syzigium chloranthum (Duth.) Merr. & Perry
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Aporusa falcifera Hk.f.
Syzygium sp.
Baccaurea bracteata M.A.
Pternandra rostrata (Cogn.) Ohwi
Aporusa falcifera Hk.f.
100
80
60
40
20
0
0
0
118
111
110
114
112
122
252
253
246
6
12
104
98
20
19
138
137
269
273
272
13
14
97
95
23
27
25
93
133
135
244
270
132
129 131
130
236
274
239
18
16
100 99
106
102
105
103
241
242 240
275
20
248
257
258
265
250 249
123
251
255
256
259
254
264
263
261 260
262
40
285
155
282
283
284
60
159
145 158
291
289
28
38
36
92
30
29
82
34
31
33
89 88
90
91
83
76
84
86
60
87
136
79 78 77
148
146
151
149
150
156
152
154 153
157 80
141
140
227
224
279
81
40
75
170
40
43
41 45
80
44
66
174
48
49
195
58
100
54
53
59
187
188
57
100
303
302
55
196
194
193
51
50
63
192
198
197
306 305
300
304
47
65
46
71
70
165
166
199
295
294
167
172
68
80
297 299
293
164
169
42
73
163
162
161
160
173
200
292
296
298
100
80
60
40
20
0
Figure 9. Distribution of Trees in Plot CNV05
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Convolvulaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Annonaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rosaceae
Polygalaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lecythidaceae
Lauraceae
Flacourtiaceae
Fagaceae
FAMILY
Anacardiaceae
Fabaceae
CNV05
42 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 43
Table 5. List of Trees in Plot CNV05
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea sp.
Swintonia glauca Engl.
Dacryodes rugosa H.J.Lam
Shorea macroptera Dyer
Baccaurea parviflora (M.A.) M.A.
Pouteria malaccensis (Clarke) Baeh.
Myristica beccarii Warb.
Shorea pinanga Scheff.
Agathis borneensis Warb.
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Swintonia glauca Engl.
Pouteria malaccensis (Clarke) Baeh.
Agathis borneensis Warb.
Shorea beccariana Burck.
Lithocarpus sp.
Memecylon myrsinoides Blume
Dead
Shorea ochracea Sym.
Garcinia dioica Bl.
Shorea pinanga Scheff.
Shorea venulosa Wood ex Meijer
Celtis philippensis Blanco
Agathis borneensis Warb.
Dead
Shorea maxwelliana King
Dead
Palaqium calophyllum (T.& B.) Pierre
Garcinia dioica Bl.
Shorea pinanga Scheff.
Scaphium macropodum (Miq.) Beumée ex Heyne
Castanopsis sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Memecylon myrsinoides Blume
Dead
Scaphium macropodum (Miq.) Beumee ex Heyne
Dead
44 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Artocarpus anisophyllus Miq.
Shorea parvifolia Dyer
Neesia altissima (Bl.) Bl.
Lophopetalum subobovatum King
Diospyros oblonga Wall ex G. Don
Pternandra rostrata (Cogn.) Ohwi
Licania splendens (Korth.) Prance
Licania splendens (Korth.) Prance
Pouteria malaccensis (Clarke) Baeh.
Sarcotheca diversifolia (Miq.) Hall.f.
Dillenia grandifolia Wall. Ex Hk.f.
Erycibe impressa Hoogl.
Shorea venulosa Wood ex Meijer
Myristica beccarii Warb.
Shorea parvifolia Dyer
Tetramerista glabra Miq.
Dacryodes incurvata (Engler) H.J. Lam
Shorea venulosa Wood ex Meijer
Agathis borneensis Warb.
Hydnocarpus kunstleri (King) Warb.
Swintonia glauca Engl.
Swintonia glauca Engl.
Shorea sp.
Pternandra rostrata (Cogn.) Ohwi
Myristica beccarii Warb.
Castanopsis sp.
Myristica beccarii Warb.
Baccaurea edulis Merr.
Baringtonia sp.
Dead
Hydnocarpus kunstleri (King) Warb.
Dead
Hydnocarpus kunstleri (King) Warb.
Scaphium macropodum (Miq.) Beumée ex Heyne
Dead
Calophyllum venulosum Miq.
Unidentified
A Field Guide to the Permanent Sample Plots • 45
Table 5. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Syzygium chloranthum (Duth.) Merr. & Perry
Shorea sp.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus kunstleri (King) Warb.
Shorea pinanga Scheff.
Dipterocarpus lowii Hk.f.
Gymnacranthera paniculata Warb.
Gluta wallichii (Hk.f.) Ding Hou
Swintonia glauca Engl.
Shorea sp.
Agathis borneensis Warb.
Shorea maxwelliana King
Koompassia malaccensis Maing. ex Benth.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Syzygium claviflorum (Roxb.) Wall. ex Cowan
Dipterocapus lowii Hk.f.
Dead
Shorea macroptera Dyer
Dead
Koompassia malaccensis Maing. ex Benth.
Garcinia dioica Bl.
Palaquium rostratum (Miq.) Burk.
Shorea seminis (de Vriese) Slooten
Dead
Shorea pinanga Scheff.
Syzygium bankense (Hassk.) Merr. & Perry
Santiria laevigata Bl. f. laevigata
Koompassia malaccensis Maing. ex Benth.
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Dead
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Agathis borneensis Warb.
Mezzetia parvifolia Becc.
Palaquium cochlariflorum V. Royen
46 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Gymnacranthera paniculata Warb.
Dead
Syzygium bankense (Hassk.) Merr. & Perry
Myristica beccarii Warb.
Shorea pinanga Scheff.
Pternandra rostrata (Cogn.) Ohwi
Shorea rubra Ashton
Horsfieldia sp.
Vatica oblongifolia Hook.f.
Xylopia malayana Hook.f. ex Th.
Shorea beccariana Scheff.
Shorea macroptera Dyer
Palaqium calophyllum (T.& B.) Pierre
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Garcinia dioica Bl.
Shorea rubra Ashton
Nephelium uncinatum Leenh
Shorea macroptera Dyer
Shorea rubra Ashton
Shorea rubra Ashton
Dead
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea rubra Ashton
Syzygium sp.
Shorea exelliptica Meijer
Artocarpus anysophyllus Miq.
Shorea maxwelliana King
Shorea parvifolia Dyer
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Diospyros sp.
Syzygium sp.
Dead
Agathis borneensis Warb.
A Field Guide to the Permanent Sample Plots • 47
Table 5. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea beccariana Burk.
Syzygium sp.
Pouteria malaccensis (Clarke) Baeh.
Xanthophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus woodii Merr.
Gymnacranthera paniculata Warb.
Garcinia nervosa Miq.
Hydnocarpus woodii Merr.
Horsfieldia sp.
Shorea rubra Ashton
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Aporusa antennifera (Airy Shaw) Airy Shaw
Nephelium uncinatum Leenh.
Shorea parvifolia Dyer
Quercus sp.
Shorea venulosa Wood ex Meijer
Dipterocapus lowii Hk.f.
Shorea rubra Ashton
Dead
Artocarpus sp.
Shorea rubra Ashton
Agathis borneensis Warb.
Diospyros sp.
Santiria griffithii (Hk.f.) Engl.
Madhuca malaccensis (Clarke) H.J.Lam.
Adinandra borneensis Kob.
Shorea rubra Ashton
Shorea maxwelliana King
Adinandra borneensis Kob.
Sindora sp.
Mangifera swintonioides Kosterm
Dead
Shorea macroptera Dyer.
Hydocarpus sp.
Madhuca malaccensis (Clarke) H.J.Lam.
Lithocarpus ewyckii Rehd.
48 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Pouteria malaccensis (Clarke) Baeh.
Shorea maxwelliana King
Shorea rubra Ashton
Koompassia malaccensis Maing. ex Benth.
Shorea rubra Ashton
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Memecylon myrsinoides Blume
Shorea venulosa Wood ex Meijer
Madhuca malaccensis (Clarke) H.J.Lam.
Lithocarpus ewyckii Rehd.
Lithocarpus ewyckii Rehd.
Artocarpus integer (Thunb.) Merr.
Elaeocarpus petiolatus (Thunb)
Dillenia grandifolia Wall.
Madhuca malaccensis (Clarke) H.J.Lam.
Garcinia dioica Bl.
Myristicaceae
Myristicaceae
Shorea rubra Ashton
Shorea venulosa Wood ex Meijer
Pouteria malaccensis (Clarke) Baeh.
Shorea rubra Ashton
Beilschmeidia sp.
Dipterocarpus lowii Hk.f.
Shorea beccariana Burk.
Garcinia sp.
Dead
Atuna excelsa (Benth.) Kosterm.
Syzygium bankense (Hassk.) Merr & Perry
Dead
Dead
Agathis borneensis Warb.
Dead
Agathis borneensis Warb.
Garcinia sp.
Anisoptera costata Korth.
Shorea venulosa Wood ex Meijer
A Field Guide to the Permanent Sample Plots • 49
Table 5. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Madhuca malaccensis (Clarke) H.J.Lam
Ternstroemia aneura Miq.
Syzygium sp.
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam
Ternstroemia aneura Miq.
Santiria griffithii (Hk.f.) Engl.
Shorea venulosa Wood ex Meijer
Baccaurea sumatrana M.A.
Swintonia glauca Engl.
Baccaurea sumatrana M.A.
Horsfieldia sp.
Syzygium jambosoides (Laut.) Merr. & Perry
Shorea exelliptica Meijer
Pouteria malaccensis (Clarke) Baeh.
Garcinia nervosa Miq.
Shorea maxwelliana King
Shorea maxwelliana King
Horsfieldia sp.
Shorea rubra Ashton
Semecarpus longifolia Bl.
Shorea maxwelliana King
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea maxwelliana King
Dead
Calophyllum lowii Hook.f.
Shorea beccariana Burk.
Gluta wallichii (Hk.f.) Ding Hou
Durio carinatus Mast.
Lithocarpus ewyckii Rehd.
Madhuca malaccensis (Clarke) H.J.Lam
Syzygium splendens (Bl.) Merr. & Perry
Gluta wallichii (Hk.f.) Ding Hou
Ixonanthes petiolaris Bl.
Dipterocarpus lowii Hk.f.
Lithocarpus ewyckii Rehd.
Shorea venulosa Wood ex Meijer
50 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea macroptera Dyer
Knema laurina (Bl.) Warb.
Adinandra borneensis Kob.
Shorea macroptera Dyer
Gluta wallichii (Hk.f.) Ding Hou
Helicia petiolaris Benn.
Syzygium jambosoides (Laut.) Merr. & Perry
Litsea firma Bl.
Gymnacranthera paniculata Warb.
Madhuca malaccensis (Clarke) H.J.Lam
Unidentified
Shorea rubra Ashton
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam.
Shorea maxwelliana King
Calophyllum lowii Hook.f.
Madhuca malaccensis (Clarke) H.J.Lam.
Calophyllum lowii Hook.f.
Shorea venulosa Wood ex Meijer
Dead
Xanthophyllum sp.
Sindora coriacea Maing. ex Prain
Melanochyla beccariana Oliv.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Swintonia glauca Engl.
Dead
Shorea rubra Ashton
Eugeissonia utilis Becc.
Gluta wallichii (Hk.f.) Ding Hou
Anisoptera sp.
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Shorea rubra Ashton
Agathis borneensis Warb.
Eugeissonia tristis Griff.
A Field Guide to the Permanent Sample Plots • 51
Table 5. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
Species
Koompassia malaccensis Maing. ex Benth.
Castanopsis sp.
Gymnacranthera paniculata Warb.
Tristaniopsis whiteana (Griff.) Wils. & Waterh.
Dead
Adinandra borneensis Kob.
Tetramerista glabra Miq.
Shorea pinanga Scheff.
Macaranga
Castanopsis
100
80
60
40
20
0
0
0
180
39
85
69
38
187
9
191
226
102
32
217
203
23
220
136
115
20
163
40
171
202
105
68
108
71
20
185
169
10
57
97
94
154
15
40
195
41
56
103
95
19
33
132
67
159
178
133
73
40
43
209
155
158
225
45
193
168
60
34
96
60
30 174
47
48
184
186
25
7
27
112
89
49
164
18
114
172
80
1
20
80
50 182
17
113
143
179
127
107
157
29
177
37
129
106
76
126
231
210
230
125
22 66 213
55
11
35
109
74
205
4
100
100
104
13
99
208
222
101
119
224
162
111
100
80
60
40
20
0
Figure 10. Distribution of Trees in Plot CNV06
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Cornaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Apocynaceae
Annonaceae
Violaceae
Ulmaceae
Theaceae
Sterculiaceae
Simarubaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Rhizophoraceae
Podocarpaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lauraceae
FAMILY
Anacardiaceae
Gnetaceae
CNV06
52 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 53
Table 6. List of Trees in Plot CNV06
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Koompassia malaccensis Maing. ex. Benth
Dead
Dipterocarpus lowii Hk.f.
Baccaurea pendula Merr.
Cephalomappa lepidota Airy Shaw
Microcos sp.
Shorea parvifolia Dyer
Dipterocarpus lowii Hk.f.
Knema cinerea (Poir.) Warb.
Licania splendens (Korth.) Prance
Syzygium sp.
Dead
Shorea parvifolia Dyer
Dead
Shorea sp.
Dipterocarpus lowii Hk.f.
Litsea castanea Hook.f.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
Dehaasia firma Bl.
Vatica rasak (Korth.) Bl.
Palaquium gutta (Hook.f.) Baill.
Tetramerista glabra Miq.
Neoscortechinia kingii (Hk.f.) Pax & Hoffm.
Tetramerista glabra Miq.
Neoscortechinia kingii (Hk.f.) Pax & Hoffm.
Shorea ovalis (Korth.) Bl.
Palaquium obtusifolium Burck.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
Amoora lanceolata Hiem.
Syzygium sp.
Lophopetalum subobovatum King
Gymnacranthera paniculata Warb.
Myristica sp.
Dacryodes incurvata (Engl.) H.J.Lam.
Elaeocarpus polystachys Wall.
54 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dehaasia firma Bl.
Koompassia malaccensis Maing. ex Benth.
Alseodaphne havilandii (Gamble) Kosterm
Scaphium macropodum (Miq.) Beumée ex Heyne
Unidentified
Gironiera nervosa Planch.
Lophopetalum subobovatum King
Dehaasia firma Bl.
Dead
Carallia brachyata (Lour.) Merr.
Shorea maxwelliana King
Memecylon myrsinoides Blume
Oncosperma horridum Scheff.
Shorea sp.
Unidentified
Artocarpus anisophyllus Miq.
Gymnacranthera forbesii (King) Warb
Mangifera magnifica Kochumen
Knema furfuracea (Hk.f. ex Th.) Warb.
Knema furfuracea (Hk.f. ex Th.) Warb.
Durio lanceolatus Mast.
Carallia brachyata (Lour.) Merr.
Dehaasia firma Kosterm.
Eugeissona utilis Becc
Lithocarpus sp.
Xylopia altissima Boerl.
Shorea parvifolia Dyer
Parastemon urophyllus DC.
Carallia brachyata (Lour) Merr.
Myristica sp.
Sarcotheca diversifolia (Miq.) Hall.f.
Calophyllum lowii Hook.f
Alstonia angustiloba Miq.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
A Field Guide to the Permanent Sample Plots • 55
Table 6. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Pternandra caerulescens Jack.
Syzygium sp.
Palaquium sumatranum Burck
Dipterocarpus lowii Hk.f.
Lophopetalum subobovatum King
Palaquium sumatranum Burck
Knema furfuracea (Hk.f.ex Th.) Warb.
Diospyros sp.
Polyosma sp.
Garcinia sp.
Eugenia chlorantum (Duth.) Merr. & Perry
Lithocarpus blumeanus (Korth.) Rehd.
Sarcotheca diversifolia (Miq.) Hall.f.
Artocarpus kemando Miq.
Tetramerista glabra Miq.
Tetramerista glabra Miq.
Unidentified
Dehaasia firma Bl.
Myristica maxima Warb.
Sindora sp.
Prunus spicata Kalkm.
Polyalthia sumatrana King
Calophyllum venulosum Zoll.
Dead
Licania splendens (Korth.) Prance
Madhuca malaccensis Lam
Baccaurea deflexa M.A.
Baccaurea angulata Merr.
Gnetum cuspidatum Bl.
Polyalthia rumphii Merr.
Eugenia fastigiata Miq.
Shorea seminis (de Vriese) Slooten
Myristica villosa Warb.
Baccaurea angulata Merr.
Eugeissona utilis Becc.
Baccaurea angulata Merr.
Madhuca sericea Lam
56 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Koompassia malaccensis Maing. ex Benth.
Sageraea elliptica (DC.) Hk.f. ex Th.
Myristica villosa Warb.
Mastixia trichotoma Bl.
Diospyros oblonga Wall. ex G.Don
Shorea ovalis (Korth.) Bl.
Myristica villosa Warb.
Baccaurea kunstleri King
Litsea laevigata Gamble
Dead
Unidentified
Unidentified
Dipterocarpus lowii Hk.f.
Dacryodes incurvata (Engler) H.J.Lam.
Hydnocarpus woodii Merr. ex Slooten
Licania splendens (Korth.) Prance
Cratoxylum formosum Benth. & Hook. f. ex Dyer
Myristica villosa Warb.
Artocarpus anisophyllus Miq.
Memecylon myrsinoides Blume
Shorea
Hydnocarpus woodii Merr. ex Slooten
Dead
Koompassia malaccensis Maing. ex Benth.
Dyera costulata Hook.f.
Shorea pauciflora King
Helicia petiolaris Benn.
Baccaurea pendulla Merr.
Melanochyla beccariana Oliv.
Dead
Rhodamnia cinerea Jack.
Artocarpus anisophyllus Miq.
Dipterocarpus lowii Hook.f.
Buchanania arborescens (Bl.) Bl.
Teysmaniodendron simplicifolium Merr.
Dead
Sarcotheca diversifolia (Miq.) Hall.f.
A Field Guide to the Permanent Sample Plots • 57
Table 6. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Syzygium chloranthum (Duth.) Merr. & Perry
Shorea parvifolia Dyer
Palaquium obtusifolium Burck.
Vatica rasak (Korth.) Bl.
Ixonanthes petiolaris Bl.
Adinandra borneensis Kob.
Vatica rasak (Korth.) Bl.
Tetramerista glabra Miq.
Melanochyla beccariana Oliv.
Koona ochracea (Elm) Merr.
Parishia maingayi Hook.f
Pternandra rostrata (Cogn.) Ohwi
Dead
Mastixia trichotoma Bl.
Durio dulcis Becc.
Lithocarpus sp.
Shorea sp.
Diospyros sp.
Dead
Durio lanceolatus Mast.
Vatica rasak (Korth.) Bl.
Dead
Myristica sp.
Palaquium rostratum (Miq.) Burck
Pternandra rostrata (Cogn.) Ohwi
Calophyllum lowii Hook.f.
Dead
Hopea sp.
Nephelium uncinatum Leenh.
Pternandra rostrata (Cogn.) Ohwi
Sarcotheca diversifolia (Miq.) Hall.f.
Myristica sp.
Tetramerista glabra Miq.
Palaquium rostratum (Miq.) Burck
Macaranga hypoleuca M.A.
Archidendron sp.
Palaquium sumatranum Burck
58 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Tetramerista glabra Miq.
Shorea parvifolia Dyer
Pternandra rostrata (Cogn.) Ohwi
Knema furfuracea (Hk.f.et Th.) Warb.
Tetramerista glabra Miq.
Palaquium rostratum (Miq.) Burck
Palaquium rostratum (Miq.) Burck
Podocarpus neriifolius D.Don.
Shorea seminis (de Vriese) Slooten
Aglaia cf. gamopetala Merr.
Celtis philippensis Blanco
Shorea parvifolia Dyer
Pternandra rostrata (Cogn.) Ohwi
Dead
Oncosperma horridum Scheff.
Adinandra borneensis Kob.
Shorea seminis (de Vriese) Slooten
Tetramerista glabra Miq.
Neesia malayana Bakh.
Gardenia tubifera Wall.
Syzygium ochneocarpa Merr.
Shorea seminis (de Vriese) Slooten
Irvingia malayana Oliv.
Shorea seminis (de Vriese) Slooten
Alstonia spathulata Blume
Canarium littorale Bl.
Lithocarpus blumeanus (Korth.) Rehd.
Palaquium rostratum (Miq.) Burck
Licania splendens (Korth.) Prance
Shorea johorensis Foxw.
Dead
Dehaasia firma Bl.
Calophyllum lowii Hook.f.
Vatica sp.
Archidendron sp.
Lophopetalum subobovatum King
Vatica sp.
A Field Guide to the Permanent Sample Plots • 59
Table 6. Continued
Tree #
223
224
225
226
227
228
229
230
231
Species
Dillenia grandifolia Wall. ex Hk.f.
Tetramerista glabra Miq.
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea sp.
Shorea sp.
Gymnacranthera contracta Warb.
Artocarpus kemando Miq.
Castanopsis motleyana King.
Lophopetalum subobovatum King
100
80
60
40
20
0
0
0
20
85
82
87
83
1
84
2
6
162
79
164
86
80
5
81
166
165
167
78
88
3
76
77
91
168
169
7
20
4
8
75
90
89 95
161
20
73
14
72
97
15
74
96
93 94
159
160
9
11
158
173
172
12
71
13
70
156
152
66
40
16
18
68
69
98
157
154 155
17
67
150
148
19
22
99
100
101
21
64
151
65
184
183
181
176
175
153
174
177
40
59
60
62
102
191
189
23
60
32
103
142
143
144
192
195
194
145
146
190
60
107
141
108
110
109
139
138
137 136
118
80
30
35
46
122
39
36 37 38
49
100
209
208
100
41
42
127
128
206
120
121
204
207
202
130
119
115 116
29
113
112
131
133
111
134
135
199
203
200 201
80
100
80
60
40
20
0
Figure 11. Distribution of Trees in Plot CNV07
Meliaceae
Lauraceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Clusiaceae
Burseraceae
Bombacaceae
Arecaceae
Apocynaceae
Annonaceae
Anacardiaceae
Verbenaceae
Tiliaceae
Thymelaeaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Rhizophoraceae
Proteaceae
Polygalaceae
Oleaceae
Olacaceae
Myrtaceae
FAMILY
Alangiaceae
Myristicaceae
CNV07
60 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 61
Table 7. List of Trees in Plot CNV07
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea sp.
Dehaasia firma Bl.
Shorea sp.
Shorea sp.
Beilschmiedia glabra Kosterm
Shorea fallax Meijer
Shorea fallax Meijer
Aglaia harmsiana Park.
Baccaurea edulis Merr.
Dead
Durio grandiflorus Kost.
Polyalthia glauca Boerl.
Garcinia maingayi Hook.f.
Spatholobus sp.
Shorea fallax Meijer
Knema furfuracea (Hk.f. ex. Th.) Warb.
Plectronia sp.
Gonystylus keithii Airy Shaw
Elateriospermum tapos Bl.
Parashorea smithiesii Wyatt-Smith ex Ashton
Tabernaemontana sphaerocarpa Bl.
Shorea ovalis (Korth.) Bl.
Artocarpus lanceifolius Roxb.
Beilschmiedia glabra Kosterm.
Polyalthia glauca Boerl.
Dead
Dead
Elateriospermum tapos Bl.
Prunus arborea (Bl.) Kalkm.
Santiria sp.
Mastixia rostrata Bl.
Ochanostachys amentacea Mast.
Aglaia ganggo Miq.
Shorea parvifolia Dyer
Baccaurea pendula Merr.
Polyalthia glauca Boerl.
Teijsmaniodendron simplicifolium Merr.
62 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Parashorea smithiesii Wyatt-Smith ex Ashton
Plectronia sp.
Litsea angulata Bl.
Pentace laxiflora Merr.
Chionanthus sp.
Dead
Teijsmaniodendron simplicifolium Merr.
Shorea ochracea Sym
Carallia brachyata (Lour.) Merr.
Shorea seminis (de Vriese) Slooten
Chisocheton divergens Bl.
Heritiera borneensis (Merr.) Kosterm.
Heritiera borneensis (Merr.) Kosterm.
Teijsmaniodendron simplicifolium Merr.
Aquilaria malaccensis Lamk.
Shorea ovalis (Korth.) Bl.
Aporusa falcifera Hk.f.
Syzygium sp.
Palaquium calophyllum Pierre ex Burck
Payena acuminta (Bl.) Pierre
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Helicia serrata (R.Br.) Bl.
Shorea fallax Meijer
Palaquium dasyphyllum Pierre
Dead
Shorea ovalis (Korth.) Bl.
Baccaurea edulis Merr.
Nephelium uncinatum Leenh
Baccaurea edulis Merr.
Shorea fallax Meijer
Dysoxylum sp.
Shorea fallax Meijer
Polyalthia sumatrana King
Artocarpus lanceifolius Roxb.
Mezzetia parvifolia Becc.
Artocarpus lanceifolius Roxb.
A Field Guide to the Permanent Sample Plots • 63
Table 7. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Shorea ovalis (Korth.) Bl.
Dacryodes rugosa H.J.Lam
Knema laurina (Bl.) Warb.
Knema furfuracea (Hk.f. ex. Th.) Warb.
Knema sp.
Diospyros sp.
Macaranga pruinosa (Miq.) Muell Arg.
Amoora lanceolata Hiern
Shorea fallax Meijer
Diospyros sp.
Shorea ovalis (Korth.) Bl.
Artocarpus lanceifolius Roxb.
Elateriospermum tapos Bl.
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Shorea fallax Meijer
Knema furfuracea (Hk.f. ex. Th.) Warb.
Knema laurina (Bl.) Warb.
Syzygium splendens (Bl.) Merr. & Perry
Palaquium rostratum Burck
Heritiera javanica (Bl.) Kosterm.
Shorea ovalis (Korth.) Bl.
Parashorea smithiesii Wyatt-Smith ex Ashton
Tetramerista glabra Miq.
Gymnacranthera paniculata Warb
Chisocheton divergens Bl.
Xylopia altissima Boerl.
Lithocarpus sundaicus (Bl.) Rehd.
Diospyros borneensis Hiern
Pometia pinnata Forst.
Shorea ovalis (Korth.) Bl.
Livistona sp.
Shorea leprosula Miq.
Madhuca oblongifolia Merr.
Parishia maingayi Hk.f.
Syzygium splendens (Bl.) Merr. & Perry
Shorea parvifolia Dyer
64 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Polyalthia glauca Boerl.
Shorea fallax Meijer
Dead
Shorea ovalis (Korth.) Bl.
Cleistanthus myrianthus (Hassk.) Krz.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Lithocarpus blumeanus (Korth.) Rehd.
Shorea parvifolia Dyer
Nephelium lappaceum Miq.
Kokoona ohracea (Elmer) Merr.
Palaquium sumatranum Burk.
Shorea ovalis (Korth.) Bl.
Xanthophyllum sp.
Livistona sp.
Koodersiodendron pinnatum (Blanco) Merr.
Adinandra sp.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pometia pinnata Forst.
Ficus glandulifera Wall.
Lithocarpus sp.
Syzygium sp.
Shorea parvifolia Dyer
Xanthophyllum amoenum Chod.
Polyalthia glauca Boerl.
Aporusa lunata (Miq.) Kurz.
Shorea ovalis (Korth.) Bl.
Myristica sp.
Dehaasia firma Bl.
Knema furfuracea (Hk.f. ex Th.) Warb.
Shorea parvifolia Dyer
Dead
Syzygium laxiflorum DC.
A Field Guide to the Permanent Sample Plots • 65
Table 7. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Dead
Pterocymbium tubulatum (Mast.) Pierre
Shorea fallax Meijer
Pentace laxiflora Merr.
Helicia petiolaris Benn.
Actinodaphne glomerata (Bl.) Nees
Koompassia malaccensis Maing. ex Benth.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Palaquium dasyphyllum Pierre
Palaquium dasyphyllum Pierre
Cleistanthus myrianthus (Hassk.) Krz.
Diospyros borneensis Hiern
Dehaasia firma Bl.
Dead
Shorea ovalis (Korth.) Bl.
Ficus globosa Bl.
Shorea leprosula Miq.
Diospyros borneensis Hiern
Cleistanthus myrianthus (Hassk.) Krz.
Syzygium sp.
Pentace laxiflora Merr.
Macaranga hosei King ex Hook.f.
Lithocarpus blumeanus (Korth.) Rehd.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Knema cinerea (Poir.) Warb.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer
Livistona sp.
Shorea parvifolia Dyer
Livistona sp.
Dead
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dye r
66 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
Species
Pternandra rostrata (Cogn.) Ohwi
Dead
Nepheium lapaceum Miq.
Syzygium sp.
Alangium sp.
Madhuca malaccensis (Clarke) H.J.Lam
Dehaasia firma Bl.
Shorea ovalis (Korth.) Bl.
Ostodes macrophylla B. ex H.
Hopea dryobalanoides Miq.
Bhesa paniculata Arn
Dead
Dead
Chisocheton divergens Bl.
Baccaurea kunstleri King.
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Polyalthiya lateriflora King
Quercus sp.
Macaranga pruinosa (Miq.) Muell Arg.
Dipterocarpus humeratus Sloot.
Calophyllum pulcherrimum Wall.
Parashorea smithiesii Wyatt-Smith ex Ashton
Calophyllum venulosum Miq.
100
80
60
40
20
0
0
230
232
228
227
231
1
118
252
2
111
3 4 5
110
7
125
127
20
12
11
115 106
8 9
209
270
13
17
18
40
15
19
94
96
100
144
20
28
286
193
31
30
27
24
87 86
88
145
194
60
60
26
25
81
146
192
34
32
33
82
83
147
84
295
76
75
41
45
46
43 44
40 39
80
47
42
38
77
299
301
51
52
55
68
71
59
61
100
64 63
65
67
69
70
160
161
159
162
66
56
57
158
163
164
157
54
156
48
100
173
172
166
168 167
169 165
171
170
73 72
155
154
151
74
149
297
298
180 177 176 175
179
178
182
174
292
150
183
35
36 37
79
148
80
291
296
185 184
294
293
186
187 188
189
289
290
190
288
191
287
285
284
85
21
90 89
91
97 93 92
99
143
140
141
142
275
278
274
138
104 98
131
133
137
14 16
105
103
130
109
102
128
129
134
136
283
281 280
279
282
139
199
201 200
202
273
267
40
196
272
198
197
269
271
268
265
266
263
264
262
258
203
204
205
207
206
122
123
224
121
112
120
256
254
212
211
261
259
260
221 223
124
119
117
225
226
215
216
20
248
247
249
246
250
251
220
236
237
239
238
240
234
233
235
245
244
243
242 241
0
100
80
60
40
20
0
Figure 12. Distribution of Trees in Plot CNV08
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Theaceae
Sterculiaceae
Simarubaceae
Sapotaceae
Sapindaceae
Rosaceae
Proteaceae
Polygalaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lauraceae
Icacinaceae
Flacourtiaceae
FAMILY
Anacardiaceae
Fagaceae
CNV08
A Field Guide to the Permanent Sample Plots • 67
68 • A Field Guide to the Permanent Sample Plots
Table 8. List of Trees on Plot# CNV08
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Artocarpus anisophyllus Miq.
Dacryodes rugosa H.J.Lam var. rugosa
Dialium trifolum de Wit
Shorea exelliptica Meijer
Knema glauca (Bl.) Warb.
Xanthophyllum rufum Bth.
Cryptocarya sp.
Knema glauca (Bl.) Warb.
Myristica tomentosa Warb.
Dead
Shorea parvifolia Dyer
Durio carinatus Mast.
Polyalthia lateriflora Kurz.
Prunus spicata Kalkm.
Shorea pauciflora King
Dipterocarpus confertus Sloot.
Cinnamomum sp.
Irvingia malayana Oliv.
Shorea macroptera Dyer
Shorea agami Ashton
Shorea macroptera Dyer
Dacryodes incurvata (Engler) H.J.Lam
Prunus sp.
Knema cinerea (Poir.) Warb.
Shorea pauciflora King
Syzygium tawahense (Korth.) Merr. & Perry
Dipterocarpus confertus Sloot.
Cryptocarya densiflora Bl.
Shorea agami Ashton
Diospyros sp.
Madhuca malaccensis (Clarke.) H.J.Lam
Quercus sp.
Myristica beccarii Warb.
Moultonianthus leembruggianus (Boerl. & Koord.) Steenis
Drypetes kikir Airy Shaw
Syzygium bankensis (Hassk.) Merr. & Perry
Licania splendens (Korth.) Prance
A Field Guide to the Permanent Sample Plots • 69
Table 8. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Shorea agami Ashton
Horsfieldia sp.
Litsea sp.
Horsfieldia sp.
Nephelium uncinatum Radlk. ex Leenh.
Endiandra coriacea Merr.
Horsfieldia sp.
Shorea beccariana Burck
Mangifera swintonioides Kosterm
Drypetes kikir Airy Shaw
Shorea rubra Ashton
Shorea rubra Ashton
Horsfieldia sp.
Shorea rubra Ashton
Shorea exelliptica Meijer
Dead
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Syzygium claviflora Wall.
Shorea exelliptica Meijer
Dead
Durio lanceolatus Mast.
Shorea maxwelliana King
Shorea maxwelliana King
Dead
Agathis borneensis Warb.
Santiria griffithii (Hk.f.) Engl.
Knema sp.
Gymnacranthera paniculata Warb
Knema sp.
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Mangifera magnifica Kochumen
Memecylon myrsinoides Bl.
Shorea beccariana Burck
Pimeleodendron papaveroides J.J.S.
Scaphium macropodum (Miq.) Beumee ex Heyne
70 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Aporusa sp.
Shorea beccariana Burck
Durio lanceolatus Mast.
Unidentified
Xylopia malayana Hk. f. Thoms.
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Canarium littorale Bl.
Pimeleodendron papaveroides J.J.S.
Shorea sp.
Dipterocarpus humeratus Sloot.
Sindora beccariana Back.
Nephelium uncinatum Radlk. ex Leenh.
Mesua cornata Stevens
Canarium littorale Bl.
Syzygium sp.
Shorea agami Ashton
Shorea beccariana Burck
Knema sp.
Shorea sp.
Shorea macroptera Dyer
Dead
Quercus subsericea A.Camus
Shorea maxwelliana King
Shorea beccariana Burck
Licania splendens (Korth.) Prance
Neesia malayana Bakh.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum pulcherrimum Wall.
Dacryodes rugosa H.J.Lam
Artocarpus odoratissimus Blanco
Pternandra galeata Ridl.
Dead
Hydnocarpus woodi Merr.
Syzygium sp.
Quercus subsericea A.Camus
A Field Guide to the Permanent Sample Plots • 71
Table 8. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Moultonianthus leembruggianus (Boerl. & Koord.) Steenis
Diospyros sp.
Prunus spicata Kalkm.
Oncosperma horridum Scheff.
Dead
Aporusa falcifera Hk.f.
Myristica sp.
Canarium denticulatum H.J.Lam
Shorea exelliptica Meijer
Oncosperma horridum Scheff.
Shorea rubra Ashton
Shorea exelliptica Meijer
Shorea maxwelliana King
Artocarpus integer (Thunb.) Merr.
Dead
Shorea beccariana Burck
Stemonurus grandifolius Becc.
Shorea rubra Ashton
Shorea maxwelliana King
Melanochyla pulvinervis (Bl.) Ding Hou
Dead
Shorea maxwelliana King
Shorea macroptera Dyer
Dead
Agathis borneensis Warb.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Lithocarpus lucidus (Roxb.) Rehd.
Dialium maingayi Backer
Shorea beccariana Burck
Elaeocarpus petiolatus Wall.
Knema sp.
Shorea maxwelliana King
Shorea parvifolia Dyer
Cinnamomum iners Reinw.
Dehaasia brachybotrys (Merr.) Kosterm.
Dipterocarpus humeratus Sloot.
72 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Dacryodes incurvata (Engler) H.J.Lam
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Dead
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Koompassia malaccensis Maing. ex Benth.
Artocarpus integer (Thunb.) Merr.
Dillenia grandifolia Wall. ex Hk.f.
Shorea maxwelliana King
Shorea rubra Ashton
Shorea exelliptica Meijer
Shorea rubra Ashton
Hydnocarpus woodii Merr.
Madhuca sp.
Quercus sp.
Shorea exelliptica Meijer
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Pouteria malaccensis (Clarke) Baeh.
Knema glauca (Bl.) Warb.
Shorea agami Ashton
Shorea seminis (de Vriese) Slooten
Cryptocarya tomentosa Bl.
Prunus spicata Kalkm.
Madhuca sericea H.J. Lam
Shorea agami Ashton
Oncosperma horridum Scheff.
Palaquium obtusifolium Burck.
Dead
Syzygium splendens (Bl.) Merr. & Perry
Tetramerista glabra Miq.
Neesia malayana Bakh.
Pouteria malaccensis (Clarke) Baeh.
A Field Guide to the Permanent Sample Plots • 73
Table 8. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Tetramerista glabra Miq.
Kokoona ochracea Merr.
Shorea pauciflora King
Syzygium splendens (Bl.) Merr. & Perry
Santiria tomentosa Bl.
Shorea beccariana Burck
Hydnocarpus sp.
Disoxylum sp.
Lithocarpus lucidus (Roxb.) Rehd.
Mangifera swintonioides Ashton
Lithocarpus ewyckii Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck
Helicia petiolaris Benn.
Shorea parvifolia Dyer
Shorea beccariana Burck
Gluta wallichii (Hk.f.) Ding Hou
Horsfieldia sp.
Shorea rubra Ashton
Shorea maxwelliana King
Shorea agami Ashton
Syzygium bankensis (Hassk,) Merr. & Perry
Dead
Dyera costulata (Miq.) Hk.f.
Dead
Shorea beccariana Burck
Horsfieldia sp.
Dead
Dead
Horsfieldia sp.
Diospyros everetii Merr.
Unidentified
Shorea rubra Ashton
Dead
Vatica umbonata (Hook.f.) Burk
Shorea rubra Ashton
Dead
74 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam
Sindora leiocarpa Backer
Shorea rubra Ashton
Shorea artinervosa Sym.
Shorea artinervosa Sym.
Glochidion
Dead
Shorea rubra Ashton
Shorea venulosa Wood. ex Meijer
Koompassia malaccensis Maing. ex Benth.
Santiria sp.
Shorea artinervosa Sym.
Dacryodes rogosa H.J.Lam
Horsfieldia sp.
Hydnocarpus woodii Merr.
Shorea maxwelliana King
Xanthophyllum
Artocarpus dadah Miq.
Shorea maxwelliana King
Vatica umbonata (Hook.f.) Burk.
Artocarpus anysophyllus Miq.
Semecarpus longifolia Bl.
Durio lanceolatus Mast.
Shorea exelliptica Meijer
Canarium litorale Bl.
Shorea beccariana Burck
Shorea exelliptica Meijer
Memecylon myrsinoides Bl.
Shorea maxwelliana King
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Dead
Shorea maxwelliana King
Dead
Shorea parvifolia Dyer
Calophyllum pulcherrimum Wall.
A Field Guide to the Permanent Sample Plots • 75
Table 8. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Horsfieldia sp.
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Prunus sp.
Gluta wallichii (Hk.f.) Ding Hou
Mezzetia parviflora Becc.
Lithocarpus sp.
Lithocarpus lucidus (Roxb.) Rehd.
Syzygium claviflora Wall.
Santiria oblongifolia Bl.
Melanochyla beccariana Oliv.
Shorea macroptera Dyer
Myristica sp.
Lithocarpus sp.
Shorea rubra Ashton
Licania splendens (Korth.) Prance
Dead
Dead
Hydnocarpus woodii Merr.
Shorea beccariana Burck
Hydnocarpus woodii Merr.
Lithocarpus sp.
Ternstroemia aneura Miq.
Mezzetia parviflora Becc.
Tetramerista glabra Miq
Knema cinerea (Poir.) Warb.
Unidentified
Prunus spicata Kalkm.
Shorea parvifolia Dyer
Knema sp.
Palaquium calophyllum Pierre ex Burck
Tetramerista glabra Miq.
Shorea macroptera Dyer
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Shorea macroptera Dyer
Dipterocarpus lowii Hook.f.
76 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
297
298
299
300
301
302
Species
Lithocarpus sp.
Tetramerista glabra Miq.
Nephelium maingayi Hiern.
Tetramerista glabra Miq.
Shorea macroptera Dyer
Unidentified
100
80
60
40
20
0
264
0
1
4
113
5
6
105
106
13
2
3
255
122
11
16
127
126 128
123
257
20
20
104
256
242
258
269
7
107
129 125
109
108
112
246
247
260
262
265
266
251
111
110
114
0
239
22
23
95
241
238
270
271
279
92
24
25
26
30
100 99 98
94
93
91
137
90
40
31
140
139
84
283
33
85
32
80
144
226
40
39
38
37
146
60
79
60
41
78
77
152
147
149
212
215 208
286
284 285
214
222 221
282
227
158
35 34
40
234
87
280
237 236
135
273
274
276
275
75
48
42
291
204
293
295
203
294
43
44
47
49
67
70
71
69
80
50
53 52
68
63
170
302
304
66
51
62
56
57
60
181
59
100
179
178
177
187
184
189
192
190
303
307 310
311
309
171
180
54
100
306 312
305
308
188
193
191
172
195
202
201 197 199
196
200
198
205
80
296 297
292
290
73
211
46
45
76
210
207
289
100
80
60
40
20
0
Figure 13. Distribution of Trees in Plot CNV09
Fagaceae
Fabaceae
Euphorbiacea
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Ulmaceae
Thymelaeaceae
Theaceae
Sterculiaceae
Sapotaceae
Rubiaceae
Rosaceae
Proteaceae
Polygalaceae
Myrtaceae
Myristicaceae
Meliaceae
Linaceae
Lauraceae
FAMILY
Anacardiaceae
Flacourtiaceae
CNV09
A Field Guide to the Permanent Sample Plots • 77
78 • A Field Guide to the Permanent Sample Plots
Table 9. List of Trees in Plot CNV09
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Shorea rubra Ashton
Dipterocarpus stellatus Vesque
Xanthophyllum parvum Chod.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Dehaasia sp.
Dead
Helicia petiolaris Benn.
Agathis borneensis Warb.
Shorea rubra Ashton
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Shorea rubra Ashton
Syzygium bankense (Hassk.) Merr.& Perry
Syzygium bankense (Hassk.) Merr.& Perry
Gluta wallichii (Hk.f.) Ding Hou
Syzygium bankense (Hassk.) Merr.& Perry
Agathis borneensis Warb.
Shorea exelliptica Meijer
Palaquium rostratum (Miq.) Burck.
Shorea beccariana Burck.
Vatica umbonata (Hook.f.) Burck.
Shorea maxwelliana King
Baccaurea racemosa M.A.
Syzygium bankense (Hassk.) Merr.& Perry
Dipterocarpus stellatus Vesque
Shorea beccariana Burck.
Shorea exelliptica Meijer
Shorea parvifolia Dyer
Palaquium callophyllum (T.& B.) Pierre
Shorea venulosa Wood. ex Meijer
Syzygium sp.
Cleistanthus sumatranus M.A.
Koompassia malaccensis Maing. ex Benth
A Field Guide to the Permanent Sample Plots • 79
Table 9. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Syzygium sp.
Shorea venulosa Wood. ex Meijer
Gonystylus forbesii Gilg.
Koompassia malaccensis Maing. ex Benth
Shorea rubra Ashton
Shorea maxwelliana King
Madhuca erythrophylla (K.ex G.) H.J.Lam
Gluta wallichii (Hk.f.) Ding Hou
Durio lanceolatus Mast.
Gymnacranthera paniculata Warb.
Knema intermedia (Bl.) Warb.
Shorea parvifolia Dyer
Dacryodes incurvata (Engler) H.J.Lam
Shorea macroptera Dyer
Agathis borneensis Warb.
Shorea beccariana Burck.
Dipterocarpus stellatus Vesque
Horsfieldia sp.
Shorea parvifolia Dyer
Hydnocarpus kunstleri (King) Warb.
Atuna excelca (Benth.) Kosterm
Dehaasia polyneura (Miq.) Kosterm
Dillenia grandifolia Wall. ex. Hk.f.
Santiria laevigata Bl.
Dacryodes rugosa (Bl.) H.J.Lam
Dipterocarpus stellatus Vesque
Madhuca malacensis (Clarke) H.J.Lam
Syzygium bankense (Hassk.) Merr.& Perry
Spatholobus sp.
Gymnacranthera paniculata Warb.
Artocarpus integer (Thumb.) Merr.
Xanthophyllum amoenum Chod.
Koompassia malaccensis Maing. ex Benth
Garcinia parvifolia Miq.
Horsfieldia sp.
Shorea macroptera Dyer
Cleistanthes myrianthus Krz.
80 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Horsfieldia glabra (Bl.) Warb,
Shorea beccariana Burck.
Palaquium rostratum (Miq.) Burck
Shorea venulosa Wood ex Meijer
Syzygium sp.
Agathis borneensis Warb.
Dead
Gonystylus forbesii Gilg.
Calophyllum pulcherrimum Wall.
Gluta wallichii (Hk.f.) Ding Hou
Cleistanthus myrianthus Krz.
Agathis borneensis Warb.
Dipterocarpus stellatus Vesque
Agathis borneensis Warb.
Horsfieldia sp.
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Sindora leiocarpa Backer
Vatica umbonata (Hook.f.) Burck.
Shorea parvifolia Dyer
Shorea macroptera Dyer
Dead
Agathis borneensis Warb.
Vatica umbonata (Hook.f.) Burck.
Hydnocarpus woodii Merr.
Dipterocarpus stellatus Vesque
Xanthophyllum
Dipterocarpus stellatus Vesque
Unidentified
Horsfieldia glabra (Bl.) Warb.
Gymnacranthera paniculata Warb.
Neonauclea excelsa (Bl.) Merr.
Shorea maxwelliana King
Madhuca oblongifolia Merr.
Dipterocarpus lowii Hk.f.
Shorea macroptera Dyer
Helicia petiolaris Benn.
A Field Guide to the Permanent Sample Plots • 81
Table 9. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Syzygium bankense (Hassk.) Merr. & Perry
Santiria griffithii (Hk.f.) Engl.
Vatica umbonata (Hook.f.) Burck.
Barringtonia scortechinii King
Mangifera magnifica Kochumen
Shorea maxwelliana King
Horsfieldia sp.
Dipterocarpus stellatus Vesque
Podocarpus neriifolius D.Don
Agathis borneensis Warb.
Scaphium macropodum (Miq.) Beumee ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus urceolaris (Jack) Merr.
Parastemon urophyllus A.DC.
Licania splendens (Korth.) Prance
Shorea beccariana Burck.
Shorea parvifolia Dyer
Parastemon urophyllus A.DC.
Palaquium cochlearifolium van Royen
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Shorea maxwelliana King
Swintonia glauca Engl.
Agathis borneensis Warb.
Shorea maxwelliana King
Scaphium macropodum (Miq.) Beumee ex Heyne
Santiria griffithii (Hk.f.) Engl.
Hydnocarpus kunstleri (King) Warb
Vatica umbonata (Hook.f.) Burck.
Agathis borneensis Warb.
Madhuca erythrophylla (K.ex G.) H.J.Lam
Calophyllum pulcherrimum Wall.
Syzygium bankense (Hassk.) Merr. & Perry
Mangifera magnifica Kochumen
Shorea maxwelliana King
Shorea rubra Ashton
82 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea venulosa Wood ex Meijer
Dead
Memecylon oleaefolium Bl.
Koompassia malaccensis Maing. ex Benth
Horsfieldia sp.
Melanochyla beccariana Oliv.
Aglaia griffithii Kurz.
Dead
Hydnocarpus kunstleri (King) Warb
Melanochyla beccariana Oliv.
Carallia brachyata (Lour.) Merr.
Sindora leiocarpa Backer.
Syzygium tawahense (Korth.) Merr. & Perry
Podocarpus neriifolius D.Don
Shorea agami Ashton
Syzygium tawahense (Korth.) Merr. & Perry
Shorea beccariana Burck.
Dead
Dacryodes rugosa (Bl.) H.J.Lam
Baccaurea kunstleri King
Palaquium cochlearifolium van Royen
Durio carinatus Mast.
Dillenia grandifolia Wall. ex Hk.f.
Litsea nidularis Gamble
Shorea beccariana Burck.
Syzygium tawahense (Korth.) Merr. & Perry
Artocarpus integer (Thumb.) Merr.
Dead
Shorea rubra Ashton
Santiria griffithii (Hk.f.) Engl.
Castanopsis ?argentea A.DC
Dipterocarpus stellatus Vesque
Baccaurea sumatrana M.A.
Shorea agami Ashton
Ternstroemia aneura Miq.
Mezzetia parvifolia Becc.
Dead
A Field Guide to the Permanent Sample Plots • 83
Table 9. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dipterocarpus stellatus Vesque
Syzygium sp.
Dipterocarpus stellatus Vesque
Parastemon urophyllus A.DC.
Myristica beccarii Warb.
Swintonia glauca Engl.
Lithocarpus sp.
Canarium litorale Bl.
Dead
Shorea exelliptica Meijer
Santiria griffithii (Hk.f.) Engl.
Shorea venulosa Wood ex Meijer
Palaquium cochlearifolium van Royen
Gymnacranthera contracta Warb.
Dacryodes incurvata (Engler) H.J.Lam
Ternstroemia aneura Miq.
Hydnocarpus kunstleri (King) Warb
Agathis borneensis Warb.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus woodii Merr.
Prunus arborea (Bl.) Kalkm.
Shorea macroptera Dyer
Agathis borneensis Warb.
Dead
Scaphium macropodum (Miq.) Beumée ex Heyne
Madhuca malaccensis (Clarke) H.J.Lam
Prunus arborea (Bl.) Kalkm.
Horsfieldia sp.
Hydnocarpus kunstleri (King) Warb
Hydnocarpus kunstleri (King) Warb
Dehaasia incrassata (Jack.)Kosterm.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Litsea firma Bl.
Unidentified
Koompassia malaccensis Maing. ex Benth
Dipterocarpus lowii Hk.f.
84 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Hydnocarpus kunstleri (King) Warb
Shorea pauciflora King
Madhuca oblongifolia Merr.
Madhuca malacensis (Clarke)H.J.Lam
Shorea macroptera Dyer
Agathis borneensis Warb.
Sindora leiocarpa Backer
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Artocarpus odoratissimus Blanco
Endiandra coriacea Merr.
Parastemon urophyllus A.DC.
Ixonanthes petiolaris Bl.
Oncosperma horridum Scheff.
Dehaasia firma Bl.
Diospyros laevigata Bakh.
Shorea rubra Ashton
Shorea beccariana Burck.
Agathis borneensis Warb.
Agathis borneensis Warb.
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Syzygium
Agathis borneensis Warb.
Agathis borneensis Warb.
Agathis borneensis Warb.
Agathis borneensis Warb.
Calophyllum pulcherrimum Wall.
Agathis borneensis Warb.
Dacryodes rostrata H.J.Lam
Dipterocarpus stellatus Vesque
Shorea parvifolia Dyer
Shorea maxwelliana King
Dead
A Field Guide to the Permanent Sample Plots • 85
Table 9. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Dipterocarpus stellatus Vesque
Callophylum venulosum Miq.
Shorea parvifolia Dyer
Dead
Gymnacranthera contracta Warb.
Santiria laevigata Bl.
Dipterocarpus stellatus Vesque
Pimeleodendron papaveroides J.J.S.
Agathis borneensis Warb.
Celtis sp.
Oncosperma horridum Scheff.
Ctenolophon parvifolius Oliv.
Dead
Melanochila elmeri Merr.
Garcinia maingayi Hook.f.
Horsfieldia glabra (Bl.) Warb.
Alstonia spathulata Blume
Eugeissona utilis Becc
Madhuca oblongifolia Merr.
Gymnacranthera paniculata Warb.
Madhuca borneensis van Royen
Dead
Shorea macroptera Dyer
Shorea parvifolia Dyer
Adinandra borneensis Kobuski
Garcinia celebica L.
Pimeleodendron papaveroides J.J.S.
Calophyllum venulosum Miq.
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Palaquium calophyllum (T. & B.) Pierre
Dipterocarpus stellatus Vesque
86 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
307
308
309
310
311
312
Species
Artocarpus integer (Thunb.) Merr.
Shorea maxwelliana King
Baccaurea angulata Merr.
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Agathis borneensis Warb.
Palaquium calophyllum (T. & B.) Pierre
Callophyllum venulosum Miq.
Syzygium bankense (Hassk.) Merr. & Perry.
Calophyllum pulcherrimum Wall.
Durio lanceolatus Mast.
Palaquium gutta (Hook.f.) Bail.
Swintonia glauca Engl.
Swintonia glauca Engl.
Hydnocarpus kunstleri (King) Warb
Lithocarpus urceolaris (Jack) Merr.
A Field Guide to the Permanent Sample Plots • 87
Table 10. List of Trees in Plot CNV10
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Artocarpus anisophyllus Miq.
Microcos cinnamomifolia Burr.
Buchanania arborescens (Bl.) Bl.
Dillenia grandifolia Wall. ex. Hk.f.
Artocarpus anisophyllus Miq.
Mezzetia parviflora Miq.
Girroniera subaequalis Planch.
Koordersiodendron pinnatum (Blanco) Merr.
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Gluta wallichii Ding Hou
Agathis borneensis Warb.
Palaquium obtusifolium Burck
Koompassia malaccensis Maing. ex Benth.
Atuna excelsa (Jack) Kosterm.
Dead
Tetramerista glabra Miq.
Koordersiodendron pinnatum (Blanco) Merr.
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
Shorea macroptera Dyer
Lithocarpus urceolaris (Jack) Merr.
Palaquium hexandrum Engl.
Syzygium sp.
Shorea exelliptica Meijer
Shorea maxwelliana King
Shorea macroptera Dyer
Hydnocarpus kunstleri (Kirng) Warb.
Baccaurea sumatrana M.A.
Shorea maxwelliana King
Baccaurea edulis Merr.
Agathis borneensis Warb.
Helicia fuscotomentosa Suesseng
Shorea beccariana Burck
Shorea exelliptica Meijer
Hydnocarpus kunstleri (Kirng) Warb.
88 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dead
Shorea exelliptica Meijer
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Dead
Shorea exelliptica Meijer
Melanochyla beccariana Oliv.
Dead
Calophyllum lowii Hook.f.
Myristica sp.
Palaquium cochlearifolium van Royen
Gluta wallichii Ding Hou
Syzygium bankense (Hassk.) Merr. & Perry
Shorea maxwelliana King
Mangifera swintonioides Kosterm.
Shorea maxwelliana King
Erycibe impressa Hoogl.
Shorea maxwelliana King
Agathis borneensis Warb.
Pimeleodendron papaveroides J.J.S.
Lithocarpus sundaicus Rehd.
Gymnacranthera contracta Warb.
Agathis borneensis Warb.
Syzygium operculata (Roxb.) Nied.
Xanthophyllum vitellinum (Bl.) D.Dietr.
Durio lanceolatus Mast.
Dillenia grandifolia Wall. ex Hk.f.
Palaquium calophyllum Pierre ex Burck
Artocarpus anysophyllus Miq.
Bhesa paniculata Arn.
Shorea exelliptica Meijer
Dialium sp.
Madhuca magnifica H.J. Lam
Palaquium cochlearifolium van Royen
Shorea exelliptica Meijer
Shorea maxwelliana King
Shorea beccariana Burck
A Field Guide to the Permanent Sample Plots • 89
Table 10. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Unidentified
Shorea beccariana Burck
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck
Lithocarpus sp.
Shorea maxwelliana King
Xanthophyllum sp.
Shorea maxwelliana King
Shorea sp.
Memecylon myrsinoides Blume
Shorea exelliptica Meijer
Swintonia glauca Engl.
Garcinia macroptera Dyer
Litsea sp.
Shorea macroptera Dyer
Myristica sp.
Calophyllum venulosum Miq.
Shorea maxwelliana King
Santiria griffithii (Hk.f.) Engl.
Shorea parvifolia Dyer
Shorea beccariana Burck
Garcinia dioica Bl.
Shorea maxwelliana King
Shorea sp.
Buchanania sessilifolia Bl.
Gluta wallichii (Hk.f.) Ding Hou
Unidentified
Palaquium cochlearifolium van Royen
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea macroptera Dyer
Erycibe impressa Hoogl.
Agathis borneensis Warb.
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea exelliptica Meijer
Syzygium bankense (Hassk.) Merr. & Perry
Shorea maxwelliana King
Shorea pinanga Scheff.
90 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Nephelium maingayi Hiern.
Terminalia sp.
Shorea macroptera Dyer
Shorea exelliptica Meijer
Dead
Xylopia malayana Hook. f. Th.
Garcinia dioica Bl.
Myristica beccarii Warb.
Gymnacranthera contracta Warb.
Anisoptera sp.
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
Shorea sp.
Gymnacranthera contracta Warb.
Terminalia sp.
Shorea sp.
Shorea parvifolia Dyer
Dead
Dead
Shorea maxwelliana King
Shorea maxwelliana King
Dead
Ixonanthes petiolaris Bl.
Aporusa antennifera (Airy Shaw) Airy Shaw
Palaquium cochlearifolium van Royen
Shorea venulosa Wood ex Meijer
Gluta wallichii Ding Hou
Garcinia dioica Bl.
Garcinia macrophylla Miq.
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Gluta wallichii Ding Hou
Helicia fuscotomentosa Suesseng
Knema latericia Elmer
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Myristica sp.
A Field Guide to the Permanent Sample Plots • 91
Table 10. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Calophyllum pulcherrimum Wall.
Shorea maxwelliana King
Garcinia dioica Bl.
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Sarcotheca diversifolia (Miq.) Hall.f.
Palaquium cochlearifolium van Royen
Pternandra rostrata (Cogn.) Ohwi
Shorea beccariana Burck
Dead
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Santiria laevigata Bl. f. laevigata
Palaquium quercifolium Burck
Gluta wallichii (Hk.f.) Ding Hou
Agathis borneensis Warb.
Calophyllum lowii Hook.f.
Dead
Shorea parvifolia Dyer
Lithocarpus sp.
Shorea venulosa Wood ex Meijer
Shorea beccariana Burck
Eugeissonia utilis Becc.
Dead
Shorea parvifolia Dyer
Lithocarpus sp.
Girroniera subaequalis Planch.
Stemonurus grandifolius Becc.
Palaquium sp.
Shorea beccariana Burck
Lithocarpus sp.
Tetramerista glabra Miq.
Shorea exelliptica Meijer
Hydnocarpus kunstleri (King) Warb.
Knema furfuracea (Hk.f. ex Th.) Warb.
Dacryodes sp.
Shorea seminis (de Vriese) Slooten
Hydnocarpus gracilis (v. Sl.) Sleum
92 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dead
Macaranga conifera (Zoll.) M.A.
Erythroxylon sp.
Syzygium sp.
Eugeissonia utilis Becc.
Pternandra rostrata (Cogn.) Ohwi
Lithocarpus sp.
Dead
Eugeissonia utilis Becc.
Lithocarpus sp.
Mallotus tiliaefolia M.A.
Adinandra borneensis Kobuski
Koompassia malaccensis Maing. ex Benth.
Unidentified
Helicia petiolaris Benn.
Gymnacranthera contracta Warb.
Dead
Dead
Hydnocarpus woodii Merr.
Calophyllum lowii Hook.f.
Shorea exelliptica Meijer
Arthocarpus anisophyllus Miq.
Shorea sp.
Palaquium cochlearifolium van Royen
Shorea beccariana Burck
Ctenolophon parvifolius Oliv.
Durio lanceolatus Mast.
Shorea beccariana Burck
Shorea agami Ashton
Lithocarpus ewyckii Rehd.
Shorea maxwelliana King
Myristica sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus lowii Hk.f.
Syzygium bankense (Hassk.) Merr. & Perry
Gluta wallichii (Hk.f.) Ding Hou
A Field Guide to the Permanent Sample Plots • 93
Table 10. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Drypetes kikir Airy Shaw
Helicia petiolaris Benn.
Lithocarpus sp.
Lithocarpus sp.
Palaquium cochlearifolium van Royen
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Koordersiodendron pinnatum (Blanco) Merr.
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Dead
Scaphium macropodum (Miq.) Beumee ex Heyne
Palaquium cochlearifolium van Royen
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Erycibe impressa Hoogl.
Hopea bracteata Burck.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Dead
Lithocarpus ewyckii Rehd.
Atuna excelsa (Jack) Kosterm.
Dead
Shorea venulosa Wood ex Meijer
Palaqium quercifolium Burk.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Litsea firma Hk. f
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
Baccaurea sumatrana M.A.
Shorea venulosa Wood ex Meijer
94 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Litsea firma Hk. f
Dead
Gymnacranthera contracta Warb.
Shorea macroptera Dyer
Prunus spicata Kalkm.
Helicia petiolaris Benn.
Shorea exelliptica Meijer
Neonauclea calycina (Bart.) Merr.
Shorea parviflora Dyer
Sarcotheca diversifolia (Miq.) Hall.f.
Scaphium macropodum (Miq.) Beumée ex Heyne
Gluta walichii (Hk.f. Ding Hou
Shorea maxwelliana King
Dipterocarpus lowii Hk.f.
Hopea bracteata Burk.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Lophopetalum beccarianum Pierre
Shorea maxwelliana King
Hydnocarpus kunstleri (King) Warb.
Hydnocarpus kunstleri (King) Warb.
Shorea exelliptica Meijer
Knema glauca (Bl.) Warb.
Dipterocarpus lowii Hk.f.
Hydnocarpus kunstleri (King) Warb.
Cryptocarya crassinervia Miq.
Shorea maxwelliana King
Shorea maxwelliana King
Shorea rubra Ashton
Shorea macroptera Dyer
Syzygium bankense (Hassk.) Merr. & Perry
Myristica sp.
Xanthophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Palaquium leiocarpum Boerl.
Shorea parvifolia Dyer
A Field Guide to the Permanent Sample Plots • 95
Table 10. Continued
Tree #
297
298
299
300
301
302
303
304
305
Species
Pternandra rostrata (Cogn.) Ohwi
Gymnacranthera contracta Warb.
Lithocarpus sericobalanus E.F. Warb.
Lithocarpus sp.
Shorea sp.
Unidentified
Shorea parvifolia Dyer
Lithocarpus pulcher (King) Markgr.
Alseodaphne havilandii (Gamble) Kosterm.
100
80
60
40
20
0
0
0
152
4
2
3
143
138
11
141
139
10
157
20
128
163
162
20
22
24
26
27
19
23
32
34
132
129
130
17
18
15
14
13
133
134
127
165
31 30
28
124
40
176
175
35
37
36
120
41
40
39
38
277
181
185
44
60
109
49
98
99
186
211
50 51
52
53
54
61
209
55
194
80
67
90
197
68
92
91
87
93
69
72
200
70
74
75
79
81
100
201
199
77
76
80
198
202
100
203
295
291
290
205
294
293
204
82
195
292
88 89
108
63
64
60
59
62
206
207
193
284
106
107
57
56
58
96
80
286
285
208
287
210
283
288
289
187
189 188
97
179
180
190
100
115
101 102
116 114
177
178
183 182
213
280
281
278
60
223
279
273
184
117
121
122
173
172
167
123
168
131
126 125
164
169 170
217
214
220
222
216 215
232
233
166
234
226
227 225 224
230
229
237
235
242 236
160
265
40
266 269 270
271
267 268
272
239
240
238
136
16
12
263
262
264
159
145 144
154
155
140
146
156
158
6
253
254
251
252
261
247 245 243
248
151
8
9
150
149
246
249
250
255
260
257
20
100
80
60
40
20
0
Figure 14. Distribution of Trees in Plot CNV11
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Dipterocarpaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Ulmaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rosaceae
Rhizophoraceae
Polygalaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
FAMILY
Anacardiaceae
Lauraceae
CNV11
96 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 97
Table 11. List of Trees in Plot CNV11
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Dead
Shorea ovalis (Korth.) Bl.
Drypetes kikir Airy Shaw
Shorea fallax Meijer
Dead
Nephelium maingayi Hiern.
Eugeissona utilis Becc
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea exelliptica Meijer
Palaquium calophyllum Pierre ex Burck
Durio lanceolatus Mast.
Melanochyla beccariana Oliv
Shorea rubra Ashton
Gymnacranthera contracta Warb.
Quercus subsericea A. Camus
Canarium sp.
Lithocarpus sundaicus Rehd.
Vatica umbonata (Hook.f.) Burck.
Santiria griffithii (Hk.f.) Engl.
Koompassia malaccensis Maing.. ex Benth.
Tetramerista glabra Miq.
Shorea macroptera Dyer
Shorea exelliptica Meijer
Lophopetalum subobovatum King
Dead
Shorea macroptera Dyer
Licania splendens (Korth.) Prance
Shorea rubra Ashton
Drypetes kikir Airy Shaw
Shorea maxwelliana King
Koompassia malaccensis Maing.. ex Benth.
Termstroemia coriacea Scheff.
Dead
Knema sp.
Shorea rubra Ashton
Swintonia glauca Engl.
Shorea parvifolia Dyer
98 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Garcinia nervosa Miq.
Hydnocarpus sp.
Shorea exelliptica Meijer
Shorea parvifolia Dyer
Swintonia glauca Engl.
Dead
Shorea rubra Ashton
Eugenia fastigiata Miq.
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata Bl.
Santiria oblongifolia Bl.
Memecylon myrsinoides Blume
Scaphium macropodum (Miq.) Beumée ex Heyne
Melanochyla cf. becariana Oliv
Syzygium tawahense (Korth.) Merr. & Perry
Shorea macroptera Dyer
Hydnocarpus kunstleri (King) Warb.
Lithocarpus sp.
Madhuca malaccensis (Clarke) H.J. Lam
Shorea parvifolia Dyer
Artocarpus integer (Thunb.) Merr.
Eugeissona utilis Becc
Eugeissona utilis Becc
Shorea maxwelliana King
Shorea maxwelliana King
Santiria griffithii (Hk.f.) Engl.
Shorea agami Ashton
Shorea maxwelliana King
Syzygium sp.
Durio lanceolatus Mast.
Syzygium sp.
Shorea rubra Ashton
Dipterocarpus lowii Hk.f.
Diospyros functiculosa Bakh.
Litsea nidularis Gamble
Lophopetalum beccarianum Pierre
Shorea maxwelliana King
A Field Guide to the Permanent Sample Plots • 99
Table 11. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
Shorea sp.
Agathis borneensis Warb.
Garcinia macrophylla Miq.
Shorea sp.
Memecylon myrsinoides Blume
Shorea sp.
Dead
Dead
Calophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus lowii Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Quercus subsericea A. Camus
Agathis borneensis Warb.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Dead
Baccaurea simatrana M.A.
Shorea sp.
Shorea maxwelliana King
Santiria griffithii (Hk.f.) Engl.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Lithocarpus sp.
Dipterocarpus lowii Hk.f.
Palaquium cochlearifolium van Royen
Shorea sp.
Agathis borneensis Warb.
Shorea sp.
100 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea sp.
Palaquium cochlearifolium van Royen
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum sp.
Dead
Dead
Shorea sp.
Shorea venulosa Wood ex Meijer
Shorea sp.
Gluta wallichii (Hk.f.) Ding Hou
Xanthophyllum sp.
Anisoptera sp.
Xanthophyllum sp.
Shorea sp.
Vatica umbonata (Hook.f.) Burck.
Swintonia glauca Engl.
Syzygium sp.
Swintonia glauca Engl.
Palaquium cochleariifolium van Royen
Dialium trifoliatum de Wit.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Gymnacranthera contracta Warb.
Dead
Shorea sp.
Shorea exelliptica Meijer
Dacryodes rugosa H.J.Lam.
Durio sp.
Dead
Shorea exelliptica Meijer
Knema cinerea (Poir.) Warb.
Shorea macroptera Dyer
Memecylon myrsinoides Blme
Dead
Durio oxleyanus Griff.
A Field Guide to the Permanent Sample Plots • 101
Table 11. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea venulosa Wood ex Meijer
Syzygium sp.
Dipterocarpus stellatus Vasque
Shorea sp.
Unidentified
Syzygium sp.
Artocarpus odoratissima Blanco
Ptychopyxis bacciformis Croiz.
Ptychopyxis bacciformis Croiz.
Gluta wallichii (Hk.f.) Ding Hou
Pternandra rostrata (Cogn.) Ohwi
Rhodamnia cinerea Jack.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Hydnocarpus woodi Merr.
Ptychopyxis bacciformis Croiz.
Calophyllum lowii Hook.f.
Sarcotheca diversifolia (Miq.) Hall .f.
Swintonia glauca Engl.
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Dead
Shorea sp.
Syzygium sp.
Dead
Syzygium sp.
Agathis borneensis Warb.
Agathis borneensis Warb.
Shorea beccariana Burck.
Lophopetalum beccarianum Pierre
Shorea sp.
Syzygium bankense (Hassk.) Merr. & Perry
Shorea parvifolia Dyer
Ixonanthes petiolaris Bl.
Artocarpus anisophyllus \Miq.
Gluta wallichii (Hk.f.) Ding Hou
102 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dyera costulata (Miq.) Hk.f.
Melanochyla beccariana Oliv.
Syzygium sp.
Dipterocarpus lowii Hk.f.
Swintonia glauca Engl.
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes rostrata H.J.Lam forma pubescens
Shorea parvifolia Dyer
Drypetes kikir Airy Shaw
Shorea ovalis (Korth.) Bl.
Mangifera swintonioides Kosterm.
Palaquium eriocalyx H.J.Lam
Shorea sp.
Shorea sp.
Shorea sp.
Shorea maxwelliana King
Dipterocarpus lowii Hk.f.
Celtis philippensis Blanco
Santiria oblongifolia Bl.
Dipterocarpus humeratus Slooten
Madhuca oblongifolia Merr.
Calophyllum venullosum Zoll.
Syzygium splendens (Bl.) Merr. & Perry
Quercus sp.
Tetramerista glabra Miq.
Knema furfuracea (Hk.f. ex Th.) Warb.
Dead
Lithocarpus sp.
Dialium platycepalum Backer
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Madhuca erythrophylla H.J.Lam
Ptychopyxis bacciformis Croiz.
Dead
Shorea pauciflora King
Dead
Knema cinerea (Poir.) Warb.
A Field Guide to the Permanent Sample Plots • 103
Table 11. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Durio grandiflorus Kost.
Shorea maxwelliana King
Heritiera javanica (Bl.) Kosterm.
Semecarpus macrophylla Merr.
Garcinia nervosa Miq.
Dead
Gymnacranthera contracta Warb.
Shorea agamii Ashton
Hydnocarpus sp.
Vatica sp.
Quercus sp.
Koompassia malaccensis Maing. ex Benth.
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Cleistanthus bakonensis Airy Shaw
Hydnocarpus sp.
Canarium littorale Bl.
Nephelium maingayi Hiern
Syzygium confertum (Korth.) Merr. & Perry
Shorea parvifolius Dyer
Shorea ovalis (Korth.) Bl.
Unidentified
Shorea ovalis (Korth.) Bl.
Calophyllum venulosum Zoll.
Phoebe laevis Kosterm.
Koompassia malaccensis Maing. ex Benth.
Shorea ovalis (Korth.) Bl.
Syzygium sp.
Dacryodes sp.
Ternstroemia aneura Miq.
Shorea pauciflora King
Gluta wallichii (Hk.f.) Ding Hou
Shorea ovalis (Korth.) Bl.
Dead
Tetramerista glabra Miq.
Pterocarpidia polyneura (Miq.) Brem.
Dead
104 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
Species
Syzygium splendens (Bl.) Merr. & Perry
Adinandra dumosa Jack.
Vatica vinosa Ashton
Tetramerista glabra Miq.
Dysoxylum
Vatica vinosa Ashton
Lithocarpus sundaicus Rehd.
Hydnocarpus
Gluta wallichii (Hk.f.) Ding Hou
Shorea pauciflora King
Cinnamomum javanicum Bl.
Shorea parvifolia Dyer
Dacryodes sp.
Shorea macroptera Dyer
Pimeleodendron papaveroides J.J.S.
Unidentified
Syzygium sp.
Nephelium uncinatum Leenh.
Vatica cf. granulata Slooten
Cleistanthes myrianthus Krz.
Macaranga pruinosa M.A.
Dacryodes rostrata H.J.Lam
Dead
Carallia brachyata (Lour.) Merr.
Pternandra rostrata (Cogn.) ohwi
Horsfieldia sp.
Shorea ochracea Sym.
Pternandra rostrata (Cogn.) ohwi
Tetramerista glabra Miq.
Ternstroemia aneura Miq.
Knema laurina (Bl.) Warb.
Tetramerista glabra Miq.
Shorea ovalis (Korth.) Bl.
Oncosperma horridum Scheff.
Myristica villosa Warb.
Syzygium sp.
100
80
60
40
20
0
0
0
1
128
4
117
116
113
114
7
110
5
139
140
141
241
20
6
111
112
8
10
11
9 12 13
14 15
16
109
104
101
100
40
98
99
157
166
19
17
24
20
23
97 96 94
91
225
172
165
86
22
85
217
31
30
27
207
206
174
29
28
32
33
37
63
58
296
295
62
61
60
44
57
183
181
190
289
45
46 47
50
48
51
52
55 53
54
56
100
187
186
188
192
193
195
49
100
298
297
291
288
292
184
185
191
199 200
196
59
182
42 43
80
34
36
41
65
66
69
180
179
70
68
35
39 38
77 78
80
73
178
198
294
205
201
203
202
213
214
204
80
215
287
285
286
67
212
176 175
75
74
71
25 79
26
81
72
173
171
209
278
216
208
277
280
282
211
163
60
21
83
161
283
281
279
219
164
162
95
93
89
88
87
160
156
158
159
276
218
169 170
224
222
223
221
60
284
274
275
220
272
273
168
167
229
267
155
154
90
153
227
151
150
233
271
268
228
269
265
40
230
266
103
102
149
145
106
105
108 107
115
238
237
231
234
236
235
232
152
142
147
146
143
239
130 131
135
242
129
118
133
132
134 136
125
124
2
123
121
120
122
119
248
249
263
260 262
251
240
252
250
253
244 243
126
127
246
245
247
254
256
259
261
20
100
80
60
40
20
0
Figure 15. Distribution of Trees in Plot CNV12
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Crypteroniaceae
Clusiaceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Verbenaceae
Tiliaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Proteaceae
Oxalidaceae
Ochnaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
Lecythidaceae
Lauraceae
Junglandaceae
FAMILY
Anacardiaceae
Icacinaceae
CNV12
A Field Guide to the Permanent Sample Plots • 105
106 • A Field Guide to the Permanent Sample Plots
Table 12. List of Trees in Plot CNV12
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Dillenia grandifolia Wall. ex Hk.f.
Teijsmaniodendron simplicifolium Merr.
Litsea grandis (Wall. ex Nees) Hook.f.
Alseodaphne falcata (Bl.) Boerl.
Dacryodes rubiginosa H.J. Lam.
Syzygium sp.
Dillenia excelsa (Jack.) Gilg.
Syzygium sp.
Dialium trifoliatum de Wit
Shorea macroptera Dyer
Knema sp.
Oncosperma horridum
Oncosperma horridum
Canarium littorale Bl.
Shorea parvifolia Dyer
Koompasia malaccensis Maing ex Benth.
Lithocarpus blumeanus Rehd.
Lithocarpus blumeanus Rehd.
Crypteronia griffithii Clarke
Knema glauca (Bl.) Warb.
Sarcotheca diversifolia (Miq.) Hall.f.
Xylopia caudata Hk.f. ex Th.
Gluta wallichii Ding Hou
Dipterocarpus stellatus Vesque
Gonocaryum impressinervium Sleum.
Dialium trifoliatum de Wit
Sindora coriacea Maingayi ex Prain
Hydnocarpus kunstleri King
Baccaurea kunstleri King
Lithocarpus blumeanus Rehd.
Gymnacranthera contracta Warb.
Madhuca malaccensis (Clarke) H.J.Lam
Swintonia glauca Engl.
Koompasia malaccensis Maing ex Benth.
Shorea beccariana Burk.
Knema glauca (Bl.) Warb.
Shorea maxwelliana King
A Field Guide to the Permanent Sample Plots • 107
Table 12. Continued
Tree #
38
39
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
65
66
67
68
69
70
71
72
73
74
75
77
Species
Syzygium chloranthum
Dialium trifoliatum de Wit
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Anisoptera costata Korth.
Elaeocarpus cf pedunculatus Wall.
Durio carinatus Mast.
Agathis borneensis Warb.
Syzygium fastigiata
Shorea macroptera Dyer
Palaquium calophyllum
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus sp.
Knema cinerea (Poir) Warb.
Barringtonia scortechinii Kurz.
Knema cinerea (Poir) Warb.
Mangifera swintonioides Kosterm.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Knema laurina (Bl.) Warb.
Dyera costulata (Miq.) Hk.f.
Shorea maxwelliana King
Helicia pescotomentosa Sueneus
Shorea macroptera Dyer
Shorea parvifolia Dyer ssp. Parvifolia
Shorea maxwelliana King
Shorea maxwelliana King
Vatica rassak (Korth.) Blume
Palaquium hexandrum
Shorea excelliptica Meijer
Knema intermedia
Shorea excelliptica Meijer
Shorea macroptera Dyer
Agathis borneensis Warb.
Dacryodes rubiginosa H.J.Lam
Lithocarpus blumeanus (Korth.) Rehd.
Diospyros sp.
108 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
78
79
80
81
83
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
112
113
114
115
116
Species
Durio lanceolata Mast.
Dipterocarpus stellatus Vesque
Garcinia macrophylla Miq.
Shorea rubra
Shorea pauciflora King
Platea sp.
Shorea parvifolia Dyer
Shorea excelliptica Meijer
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Hydnocarpus woodii Merr.
Shorea maxwelliana King
Aporosa falcifera Hk.f.
Shorea maxwelliana King
Shorea rubra
Gymnacranthera contracta Warb.
Palaquium rostratum (Miq.) Burck.
Shorea parvifolia Dyer
Hydnocarpus woodii Merr.
Anisoptera
Syzygium opaca
Shorea parvifolia Dyer ssp. Parvifolia
Hydnocarpus woodii Merr.
Oncosperma horidum
Timonius wallichianus
Payena acuminatissima
Sarcotheca diversifolia (Miq.) Hall.f.
Oncosperma horidum
Oncosperma horidum
Hydnocarpus woodii Merr.
Licania splendens (Korth.) Prance
Parashorea smythiesii Wyatt-Smith ex Ashton
Neonauclea excelsa (Bl.) Merr.
Durio
Knema laurina (Bl.) Warb.
Vatica sp.
Shorea parvifolia Dyer ssp. Parvifolia
A Field Guide to the Permanent Sample Plots • 109
Table 12. Continued
Tree #
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
149
150
151
152
153
154
Species
Diospyros sp.
Knema laurina (Bl.) Warb.
Ryparosa hulletii King
Parashorea smythiesii Wyatt-Smith ex Ashton
Knema cinerea (Poir.) Warb.
Hydnocarpus woodii Merr.
Pentace discolor Merr.
Teijsmaniodendron simplicifolium Merr.
Nephelium sp.
Moultonianthus leemburgianus
Oncosperma horidum
Lithocarpus sp.
Madhuca oblongifolia
Macaranga gigantea
Madhuca oblongifolia
Lithocarpus urceolatus
Aporosa sp.
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
dead
dead
Hydnocarpus woodii Merr.
Paratocarpus venenosa Becc.
Shorea maxwelliana King
Shorea agamii Ashton
Ctenolophon parvifolius Oliv.
dead
Shorea rubra
Lithocarpus sp.
Shorea maxwelliana King
Koompasia malaccensis Maing ex Benth.
Dacryodes incurvata H.J.Lam.
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Gardenia tubifera Wall.
Shorea ovalis (Korth.) Bl.
110 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
178
179
180
181
182
183
184
185
186
187
188
189
190
191
192
Species
Palaquium rostratum (Miq.) Burck.
Shorea beccariana Burk.
Vatica umbonata (Hook.f.) Burk.
Shorea macroptera Dyer
Melanochyla beccariana Oliv.
Dipterocarpus stellatus Vesque
Agathis boneensis Warb.
Shorea macroptera Dyer
Agathis boneensis Warb.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Agathis boneensis Warb.
Gymnacranthera contracta Warb.
Shorea macroptera Dyer
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea beccariana Burk.
Sarcotheca diversifolia (Miq.) Hall.f.
Baccaurea sumatrana M.A.
Knema glauca (Bl.) Warb.
Herritiera javanica (Bl.) Kosterm
Calophyllum lowii Hook.f.
Baccaurea sumatrana M.A.
Prunus spicata Kalkm
Pternandra rostrata (Cogn.) Ohwi
Payena endertii H.J.Lam.
Drypetes kikir Airy Shaw
Shorea parvifolia Dyer
Parkia sp.
Gymnacranthera contracta Warb.
Elateriospermum tapos Bl.
Hydnocarpus kunstlerii (King) Warb.
Dialium trifoliatum de Wit
Knema cinerea (Poir.) Warb.
Castanopsis javanica (Bl.) A.Dc.
Elateriospermum tapos Bl.
Shorea agamii Ashton
Shorea parvifolia Dyer ssp. Parvifolia
A Field Guide to the Permanent Sample Plots • 111
Table 12. Continued
Tree #
193
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
211
212
213
214
215
216
217
218
219
220
221
222
223
224
225
227
228
229
230
231
232
Species
Shorea macroptera Dyer
Artocarpus anisophyllus Miq.
Artocarpus anisophyllus Miq.
Diospyros sp.
Cephalomappa lepidotula Airy Shaw
Knema cinerea (Poir.) Warb.
Artocarpus anisophyllus Miq.
Englhardia serrata Bl.
Shorea parvifolia Dyer ssp. Parvifolia
Syzygium tawahense
Artocarpus integra (Thunb.) Merr.
Artocarpus anisophyllus Miq.
Knema cinerea (Poir.) Warb.
Shorea sp.
Licania splendens (Korth.) Prance
Litsea nidularis Gamble
Cleistanthus myrianthus Krz.
Dyera costulata (Miq.) Hk.f.
Polyanthia glauca Boerl.
Artocarpus anisophyllus Miq.
Shorea rubra
Palquium sumatranum Burck.
Shorea beccariana Burk.
Syzygium sp.
Knema conferta (King.) Warb.
Knema conferta (King.) Warb.
Sarcotheca diversifolia (Miq.) Hall.f.
Syzygium perpuncticulatum
Syzygium perpuncticulatum
Sarcotheca diversifolia (Miq.) Hall.f.
Agathis boneensis Warb.
Oncosperma horridum
Pterocymbium tubulatum (Most.) Pierre
Mangifera swintonioides Kosterm.
Pimeleodendron papaveroides J.J.S.
Lithocarpus blumeanus (Korth.) Rehd.
Lithocarpus blumeanus (Korth.) Rehd.
112 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
256
259
260
261
262
263
264
265
266
267
268
269
271
272
273
Species
Sindora beccariana Back.
Teijsmaniodendron simplicifolium Merr.
Shorea leprosula Miq.
Ctenolophon parvifolius Oliv.
Palaqium gutta (Hook.f.) Baill.
Garcinia gaudichaudii
Shorea beccariana Burk.
Garcinia macrophylla Miq.
Shorea maxwelliana Miq.
Shorea parvifolia Dyer.
Diospyros sp.
Gluta wallichii (Hk.f.) Ding Hou
Licania splendens (Korth.) Prance
Baccaurea
Palaquium rostratum
Helicia pescotomentosa Sueneus
Madhuca sp.
Garcinia dioica Bl.
Shorea parvifolia Dyer.
Licania splendens (Korth.) Prance
Calophyllum pulcherrianum Wall. ex Choisy
Hydnocarpus kunstleri (King.) Warb.
Syzygium opaca
Santiria tomentosa
Parinari sp.
Sagera elliptica (DC.) Hk.f. ex Th.
Syzygium sp.
Alstonia soathulata Blume
Elateriospermum tapos Bl.
Elateriospermum tapos Bl.
Calophyllum cf. soulatri Burm.f.
Garcinia nervosa Miq.
Dehaasia firma Bl.
Koompasia malaccensis Maing ex Benth.
Parishia maingayi Hk.f.
Gymnacranthera paniculata Warb.
Anisoptera
A Field Guide to the Permanent Sample Plots • 113
Table 12. Continued
Tree #
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
291
292
293
294
295
296
297
298
Species
Vatica rassak (Korth.) Blume
Syzygium endertii
Dacryodes rostrata H.J.Lam.
Baccaurea edulis Merr.
Lithocarpus sp.
Mallotus echinatus
Hydnocarpus woodii Merr.
Dacryodes rostrata H.J.Lam.
Madhuca corthalsii (Pierre) H.J.Lam
Mangifera swintonioides Kosterm.
Baccaurea edulis Merr.
Brackenridgea palustris Bartell.
Baccaurea motleyana M.A.
Cleistanthus myrianthus Krz.
Knema laurina (Bl.) Warb.
Garcinia dioica Bl.
Vatica rassak (Korth.) Blume
Artocarpus anisophyllus Miq.
Pentace laxiflora
Sterculia rubiginosa Vent.
Dyera costulata (Miq.) Hk.f.
Licania splendens (Korth.) Prance
Shorea pinanga Scheff.
Calophyllum pulcherrimum Wall. ex Choisy
The Center for International Forestry Research (CIFOR) is a leading international
forestry research organization established in 1993 in response to global concerns about the
social, environmental, and economic consequences of forest loss and degradation. CIFOR
is dedicated to developing policies and technologies for sustainable use and management
of forests, and for enhancing the well-being of people in developing countries who rely on
tropical forests for their livelihoods. CIFOR is one of the 15 Future Harvest centres of the
Consultative Group on International Agricultural Research (CGIAR). With headquarters
in Bogor, Indonesia, CIFOR has regional offices in Brazil, Burkina Faso, Cameroon and
Zimbabwe, and it works in over 30 other countries around the world.
Donors
The Center for International Forestry Research (CIFOR) receives its major funding from
governments, international development organizations, private foundations and regional
organizations. In 2005, CIFOR received financial support from Australia,Asian Development
Bank (ADB), Belgium, Brazil, Canada, China, Centre de coopération internationale
en recherche agronomique pour le développement (CIRAD), Cordaid, Conservation
International Foundation (CIF), European Commission, Finland, Food and Agriculture
Organization of the United Nations (FAO), Ford Foundation, France, German Agency for
Technical Cooperation (GTZ), German Federal Ministry for Economic Cooperation and
Development (BMZ), Indonesia, International Development Research Centre (IDRC),
International Fund for Agricultural Development (IFAD), International Tropical Timber
Organization (ITTO), Israel, Italy, The World Conservation Union (IUCN), Japan, Korea,
Netherlands, Norway, Netherlands Development Organization, Overseas Development
Institute (ODI), Peruvian Secretariat for International Cooperation (RSCI), Philippines,
Spain, Sweden, Swedish University of Agricultural Sciences (SLU), Switzerland, Swiss Agency
for the Environment, Forests and Landscape, The Overbrook Foundation, The Nature
Conservancy (TNC), Tropical Forest Foundation, Tropenbos International, United States,
United Kingdom, United Nations Environment Programme (UNEP), World Bank, World
Resources Institute (WRI) and World Wildlife Fund (WWF).
The CIFOR Permanent Sample Plots (PSP) were
established in 1998 in conjunction with the Phase I of the
ITTO Project dealing with sustainable forest management,
integrated land use, reduced-impact logging, biodiversity
in the production forest, and cultural and social aspects
of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes,
long term growth and yield and to provide critical data for
evaluation of ecological models. For silvicultural purposes
PSP supply data on diameter and volume increment as
well as stand structure dynamics.
In particular, the PSP in the Malinau Research Forest
provides a valuable insight into the diversity of trees found
there and the growth of the forest. This guidebook has
been designed to facilitate the growing interests and needs
of visitors to the Malinau Research Forest (MRF).
PATEN MA LINAU
KAB U
I N TI M U N G