1. No category

A Field Guide
To the Permanent Sample Plots
in the Reduced-Impact Logging Block 27
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
A Field Guide
To the Permanent Sample Plots
in the Reduced-Impact Logging Block 27
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
National Library of Indonesia Cataloging-in-Publication Data
Kartawinata, Kuswata et al.
A field guide to the permanent sample plots in the reduced-impact blocks 27 at CIFOR Malinau
research forest East Kalimantan/by Kuswata Kartawinata, Hari Priyadi, Douglas Sheil, Soedarsono
Riswan, Plinio Sist, Machfudh. Bogor, Indonesia: Center for International Forestry Research (CIFOR),
2006.
131p.
ISBN: 979-24-4630-3
CAB Int. thesaurus: 1. logging 2. reduced-impact logging 3. sample plot technique 4. stand density 5.
forest trees 6. identification 7. Indonesia 8. guidelines
© 2006 by CIFOR and ITTO
All rights reserved. Published in 2006
Printed by Intiprima Karya
Cover photos by Hari Priyadi and Ahmad Zakaria
Cover design by Eko Prianto
Layout by Catur Wahyu
Maps by Mohammad Agus Salim
Published by
Center for International Forestry Research
Mailing address: P.O. Box 6596 JKPWB, Jakarta 10065, Indonesia
Office address: Jl. CIFOR, Situ Gede, Sindang Barang,
Bogor Barat 16680, Indonesia
Tel.: +62 (251) 622622; Fax: +62 (251) 622100
E-mail: [email protected]
Web site: http://www.cifor.cgiar.org
3
Contents
Foreword
4
Introduction
5
The Plot
8
Access to a Plot
9
Plant Identification
10
Distribution of PSPS
11
Plot# RIL01
12
Plot# RIL02
21
Plot# RIL03
30
Plot# RIL04
52
Plot# RIL05
60
Plot# RIL06
70
Plot# RIL07
78
Plot# RIL08
87
Plot# RIL09
96
Plot# RIL10
105
Plot# RIL11
114
Plot# RIL12
123
4
Foreword
The CIFOR Permanent Sample Plots (PSP) were established in 1998
in conjunction with the Phase 1 of the ITTO Project dealing with
sustainable forest management, integrated land use, reduced-impact
logging, biodiversity in the production forest, and cultural and social
aspects of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes, long term
growth and yield and to provide critical data for evaluation of ecological
model. For silvicultural purposes PSP supply data on, diameter and
volume increment as well as stand structure dynamics.
I therefore appreciate those who have given strong initiative and
support in the establishment of PSP in Malinau Research Forest and
continuously measure those plots. My utmost appreciation goes also
to our partners: PT INHUTANI II for logging activities, BIOMA, the
Malinau District Office for administrative research arrangements and
the Indonesian Institute of Sciences (LIPI) for scientific identification
of trees I also wish to express my sincere thanks to Dr. Kuswata
Kartawinata, Dr. Plinio Sist of Cirad Foret, Dr. Kade Sidiyasa of
FORDA and Hari Priyadi who maintain their continuing interest by
revisiting the plots and confirming the accuracy of the identity of trees
found in the plots.
Dr Kartawinata and I take the initiative to publish the guidebook to
satisfy the growing interests and needs of visitors visiting our Seturan
camp. More and more people think highly of the importance of these
plots to understand the secret of forest growth and the diversity of
trees occurring inside the permanent plots.
This initiative is not possible without continuous support of ITTO in
providing fund for both research and publication of this guide. It is
my sincerest hope that this little guide will benefit any one who is
concerned with the future of the tropical forest.
Bogor, May 2006
Dr. Petrus Gunarso
Malinau Research Forest Coordinator
5
Introduction
A set of Permanent Sample Plots (PSP) was established in the lowland
mixed dipetrocarp forest within PT Inhutani II concession at Malinau
(2º45’ – 3º15’N, 116º30’), East Kalimantan (Figure 1). They were
designed for comparative studies of conventional logging (CNV) and
reduced-impact logging (RIL) techniques under the ITTO Project PD
12/97 Rev.1 (F), Forest, Science and Sustainability: Bulungan Model
Forest. The Malinau concession is located about 30 km to the East
of CIFOR’s Bulungan Research Forest in a forest area with elevation
between 100 m to 300 m above sea level. The terrain is undulating
with slopes ranging from 10% to 70%. The mean annual rainfall in the
area is 3.790 mm, with the number of wet months of more than 9
months per year.
This pocket guideline is intended to give visitors to PSPs information
on how to access to the location of plots and distribution of trees with
dbh ≥ 20cm in each plot. The CIFOR’s Seturan station is used as a
starting point to travel to the plots.
6
Figure 1. Malinau Research Forest Map
Figure 2. Location of Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 7
8
The Plots
The objective of setting up the permanent sample plots is to monitor
long-term impact of logging on forest dynamic, growth and yield.
The plot locations were selected randomly according to the density
of harvestable timber trees (diameter at breast height or dbh ≥ 60cm)
prior to logging using the tree maps and inventory data in the logging
compartments prepared by CIFOR and PT INHUTANI II. A field
checking was carried out before delimiting each plot to asses the
efective density of harvestable trees. Between March – May 1999, 12
one-hectare plots were set up in Petak 27 (Figure 2).
Each one-hectare plot (100 m x 100 m) was horizontally corrected
during its establishment. The plot was divided into 25 subsquares
(20m x 20m), each being divided into 4 elementary squares (a,b,c,d,
in Figure 3 below). At 20 m intervals throughout the plot, PVC piping
was driven into the soil, and a metal tag attached to indicate the sub
square’s coordinates within the plot. A total of 36 PVC stakes were
placed throughout the plot. A wood panel indicating the block and
the number of the plot marks the centre of each. Three control plots
(RIL04, RIL08, RIL12) were designated and in each one, a 50 m buffer
zone was set up to avoid edge effect caused by logging activities.
After logging, all plots were re-grided (every subsquare 20m x 20 m
materialised by rope) and permanently marked in the field with iron
wood/ulin poles located at each corner.
9
Access to a Plot
Visitors are advised to refer to the map of distribution of the permanent
sample plots (Figure 4) to get an overview on how the plots are spatially
distributed. The plots are grouped based on their route. The routes
follow the best walking way to go to each plot, that is the available
skidding road in the field.
To enter a route, visitors should find direction sign in the field posted
on the entrance of each route and follow a skidding road directed by
the sign. Zoomed maps of the routes are given in Figures 4.
Figure 3. Permanent Sample Plot structure (PVC piping is located at
each number)
20m
31 0
20m
20m
c
a
b
a
b
c
d
c
c
a
b
d
c
b
a
b
a
26
a
b
a
b
c
d
c
b
a
13
a
b
d
c
d
a
b
d
23
23
24
b
a
b
d
c
d
c
d
c
b
a
b
a
b
a
b
d
c
d
b
a
16
17
9
c
b
a
18
10
11
10
d
11
d
c
b
a
3
3 0
17
16
2
East
a
12
4
4 0
5
5 0
20m
b
1
South
b
30
22
9
c
West
a
15
d
North
b
15
8
2 0
d
a
8
1 0
c
29
c
14
7
a
d
d
14
c
c
20m
c
7 0
b
d
29
22
d
d
20m
c
a
a
21
d
13 0
b
28
21
20
c
a
28
20
19 0
c
27
19
36
d
27
26
a
35
d
20m
c
25
20m
34
d
20m
d
25 0
20m
33
32
c
b
6 0
10
Plant Identification
Trees with dbh ≥ 20 cm and dbh ≥ 10 cm were identified in the field by
botanists of the Herbarium Bogoriense, Puslitbang Biologi-LIPI, Bogor.
Voucher specimens were collected for further identification and are
stored at the Herbarium Bogoriense. The identification is not final,
hence any corrections are welcome and will be very much appreciated.
The nomenclature follows T.C. Whitmore, I.G.M. Tantra and U. Sutisna
(Editor), Tree Flora of Indonesia: Check List for Kalimantan, Agency
for Forestry Research and Development, Ministry of Forestry, Bogor,
and C.G. G.J. Van Steenis, W.J.O. De Wilde, C. Kalkman, etc. (Editors),
Flora Malesiana Serie I, Volumes 1-15, 1948-2001. Note that Plot RIL03
has 800 (760 living and 40 dead) trees with dbh ≥ 10 and 221 species in
one hectare, the highest number of species and density ever recorded
in a lowland forest in Kalimantan and perhaps in the world.
an
tur
Se
CIFOR RESEARCH STATION
TPN 1
FIRE TOWER
BRIDGE
PSP
RIVER
ROAD
Legend
Route# 1
Route# 2
Route# 3
Route# 4
RIL07
RIL09
RIL10
RIL02
RIL12
RIL11
RIL06
RIL04
RIL08
RIL07, RIL11, RIL02
RIL01, RIL04
RIL09, RIL06, RIL10, RIL12
RIL05, RIL03, RIL08
RIL01
Figure 4. Distribution of Permanent Sample Plots in RIL Block (Petak 27)
RIL05
RIL03
A Field Guide to the Permanent Sample Plots • 11
100
80
60
40
20
0
0
0
1
2
111
108
104
4
3
109
6
118
116
232
237
105
113
114
236
235
106
107
112
115
234
239
5
119
117
101
120
7
121
20
128
102
8
9
124
10
125
244
13
11
95
97 96
98
222
224
99
100
123
229 228
230
225
227
242
226
231
241
103
240
20
17
127
126
40
247
250
251
216
259
15
92
81
82
86
138
16
18
23
21
24
25
77
60
27
26
83
84
85
137 140 141
79 80
91
93
94
78
19 20 22
90
89
87
88
129
135
136
134
150
211
204
147
149
30
154
151
72
70
68 67
162
28
31
35
33
66
36
34
64
80
37
58
164
163
166
167
190
38
49
50
56
51
43
47
48
40
52
60
41
42
39
55
57
177
179
176
174
180
173
181
183
182
277
278
100
46
45
44
59 54 53
165
175
191
100
276
275
178
184
274
273
192
270
170
61
169
171
193
188
271
168
194
80
62
153
152
65
32
71
74
75 76
63
29
73
69
143
202
268 269
201
199 196 195
209
206
264
155
159
146 157
161
145
160
158
144 156
148
214
263
262
215
210
261
260
213 212
131 132
133
130
221
220
60
254
252
256
255
257 258
253
249
217
218
219
245
246
40
100
80
60
40
20
0
Figure 5. Distribution of Trees in Plot RIL01
Fagaceae
Fabaceae
Euphorbiacea
Erythroxylaceae
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Ulmaceae
Thymelaeaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Polygalaceae
Podocarpaceae
Oxalidaceae
Olacaceae
Ochnaceae
Myrtaceae
Myristicaceae
Meliaceae
Lecythidaceae
Lauraceae
FAMILY
Anacardiaceae
Flacourtiaceae
RIL01
12 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 13
Table 1. List of Trees in Plot RIL01
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Aglaia cordata Hiern.
Polyalthia
Canarium littorale Bl.
Madhuca magnifica Hk.f.
Atuna sp.1
Cleistanthus myrianthus Kurz.
Kibatalia arborea (Bl.) G.Don
Tetramerista glabra Miq.
Madhuca magnifica H.J.Lam
Mangifera swintonioides Kosterm.
Atuna excelsa (Benth.) Kosterm.
Ochanostachys amentacea Mast.
Ochanostachys amentacea Mast.
Dead
Buchanania arborescens (Bl.) Bl.
Shorea pauciflora King
Santiria malaccensis Pierre
Madhuca magnifica H.J.Lam
Shorea macroptera Dyer
Pouteria malaccensis Pierre
Endiandra rubescens Miq.
Artocarpus anisophyllus Miq.
Neesia synandra Mast.
Vatica maingayi Dyer
Santiria griffithii (Hk.f.) Engl.
Shorea macroptera Dyer
Knema cinerea Warb.
Shorea parvifolia Dyer ssp. parvifolia
Koompassia malaccensis Maing ex Benth.
Shorea parvifolia Dyer ssp. parvifolia
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes rostrata (Bl.) Lam
Eugenia sp.
Shorea parvifolia Dyer ssp. parvifolia
Madhuca malaccensis (Clarke) H.J.Lam
Lithocarpus cf. ewickii (Korth.) A. Kamus
Scaphium macropodum (Miq.) Beumée ex Heyne
14 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Canarium littorale Bl.
Dipterocarpus confertus Sloot.
Phoebe laevis Kosterm
Madhuca magnifica H.J.Lam
Hydnocarpus kunstleri (King) Warb.
Syzygium rostratum (Blume) DC.
Nauclea subdita (Korth.) Steud
Xanthophyllum rufum Bth.
Artocarpus kemando Miq.
Knema lunduensis (Warb.) de Wilde
Barringtonia macrostachya Kurz
Artocarpus sp.
Atuna excelsa sp.1
Pimeleodendron papaveroides J.J.S.
Shorea excelliptica Meijer
Gluta wallichii (Hk.f.) Ding Hou
Sarcotheca diversifolia (Miq.) Hall.f.
Hydnocarpus woodii Merr.
Shorea artinervosa Sym
Shorea brunnescens Ashton
Gomphia serrata (Gaertn.) Kanis
Hydnocarpus kunstleri (King) Warb.
Mangifera swintonioides Kosterm.
Durio malaccensis Mast.
Gluta wallichii (Hk.f.) Ding Hou
Oncosperma horridum Scheff.
Oncosperma horridum
Shorea pinanga Scheff.
Euggesona utilis Becc.
Nauclea sp.
Shorea pinanga Scheff.
Vatica umbonata (Hook.f.) Burck.
Shorea macroptera Dyer
Syzygium confertum (Korth.) Merr & Perry
Scaphium macropodum (Miq.) Beumee ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
A Field Guide to the Permanent Sample Plots • 15
Table 1. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Vatica umbonata (Hook.f.) Burck.
Dialium sp.
Garcinia dioica Bl.
Scaphium macropodum (Miq.) Beumee ex Heyne
Vatica umbonata (Hook.f.) Burck.
Mangifera swintonioides Kosterm.
Vatica umbonata (Hook.f.) Burck.
Shorea exelliptica Meijer
Artocarpus odoratisima Blanco
Myristica villosa Warb.
Myristica beccarii Warb.
Artocarpus lanceifolius Roxb.
Pimeleodendron papaveroides J.J.S.
Madhuca magnifica H.J.Lam
Shorea artinervosa Sym
Madhuca magnifica H.J.Lam
Dialium sp.
Shorea brunnescens Ashton
Gluta wallichii (Hk.f.) Ding Hou
Shorea artinervosa Sym
Castanopsis tungurut DC
Santiria griffithii (Hk.f.) Engl.
Bhesa paniculata Arn.
Litsea mappacea (Bl.) Boerl.
Scaphium macropodum (Miq.) Beumee ex Heyne
Lithocarpus urceolaris (Jack.) Merr.
Prainea limpato (Miq.) Heyne
Madhuca magnifica H.J.Lam
Dillenia reticulata King
Shorea fallax Meijer
Macaranga conifera Muel Arg.
Knema sp.
Lithocarpus urceolaris (Jack.) Merr.
Macaranga conofera Muel Arg.
Erythroxylon cuneatum (Miq.) Kurz.
Dead
Koompassia malaccensis Maing ex Benth.
16 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea parvifolia Dyer ssp. parvifolia
Lithocarpus sp.1
Shorea macroptera Dyer
Shorea fallax Meijer
Tetramerista glabra Miq.
Nephelium maingayi Hiern
Durio dulcis Becc.
Campnosperma auricululatum (Bl.) Hk.f.
Santiria laevigata Bl.
Dillenia grandifolia Wall. ex Hk.f.
Baccaurea cf. simatrana M.A.
Gironniera hirta Radlk.
Dacryodes rostrata (Bl.) H.J.Lam
Diospyros puncticulosa Bakh.
Dialium maingayi Backer
Madhuca malaccensis (Clarke) H.J.Lam
Shorea macroptera Dyer
Shorea parvifolia Dyer ssp. parvifolia
Castanopsis oviformis Soepadmo
Shorea beccariana Burck.
Palaquium ferox H.J.Lam
Plectronia sp.
Dipterocarpus confertus Sloot.
Shorea beccariana Burck.
Dyera costulata (Miq.) Hk.f.
Vatica sp.
Shorea beccariana Burck.
Hydnocarpus woodii Merr.
Shorea macroptera Dyer
Shorea beccariana Burck.
Shorea parvifolia Dyer ssp. parvifolia
Shorea parvifolia Dyer ssp. parvifolia
Shorea beccariana Burck.
Quercus sp.
Shorea beccariana Burck.
Shorea beccariana Burck.
Shorea beccariana Burck.
A Field Guide to the Permanent Sample Plots • 17
Table 1. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea venulosa Wood ex Meijer
Shorea beccariana Burck.
Ternstroemia aneura Miq.
Calophyllum pulcherrimum Wall. ex Choisy.
Hydnocarpus kunstleri (King) Warb.
Mangifera swintonioides Kosterm.
Agathis borneensis Warb.
Gonystylus forbesii Gilg.
Shorea rubra Ashton
Myristica maxima Warb.
Hydnocarpus kunstleri (King) Warb.
Hydnocarpus kunstleri (King) Warb.
Canarium littorale Bl.
Shorea beccariana Burck.
Hydnocarpus kunstleri (King) Warb.
Scaphium macropodum (Miq.) Beumee ex Heyne
Durio lanceolatus Mast.
Shorea maxwelliana King
Artocarpus anisophyllus Miq.
Gluta wallichii (Hk.f.) Ding Hou
Eugenia attenuatum (miq.) Merr & Perry
Calophyllum pulcherrimum Wall. ex Choisy.
Shorea venulosa Wood ex Meijer
Dead
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Shorea exelliptica Meijer
Myristica maxima Warb.
Shorea macroptera Dyer
Cratoxylum arborescens (Vahl.) Bl.
Shorea pinanga Scheff.
Licania splendens (Korth.) Prance
Gluta wallichii (Hk.f.) Ding Hou
Santiria griffithii (Hk.f.) Engl.
Blumeodendron elateriospermum J.J.Sm.
Shorea agami Ashton
18 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Shorea beccariana Burck.
Lithocarpus cantleyanus
Scaphium macropodum (Miq.) Beumee ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Garcinia dioica Bl.
Shorea agami Ashton
Calophyllum lowii Hook.f.
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Mangifera swintonioides Kosterm.
Cleisthanthus sp.1
Unidentified
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum lowii Hook.f.
Gironniera subaequalis Planch.
Scaphium macropodum (Miq.) Beumee ex Heyne
Calophyllum sp.
Licania splendens (Korth.) Prance
Palaquium cochlearifolium van Royen
Shorea macroptera Dyer
Sindora beccariana Backer ex de Wit
Dipterocarpus lowii Hk.f.
Dead
Quercus gmelifflora Bl.
Scaphium macropodum (Miq.) Beumee ex Heyne
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Artocarpus integer (Thunb.) Merr.
Shorea beccariana Burck.
Endiandra rubescens Miq.
Shorea beccariana Burck.
Shorea pinanga Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Shorea parvifolia Dyer
Artocarpus odoratissimus Blanco
Shorea macroptera Dyer
A Field Guide to the Permanent Sample Plots • 19
Table 1. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Dead
Dacryodes rubiginosa (Benn.) H.J.Lam
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Knema cinerea Warb.
Oncosperma horridum Scheff.
Gonystylus cf. forbesii Gilg.
Shorea parvifolia Dyer
Knema latericia Elmer
Shorea macroptera Dyer
Nephelium cf. cuspidatum Bl.
Dead
Adinandra sarosanthera Miq.
Shorea fallax Meijer
Cleisthanthus myrianthus Krz.
Palaquium ferox H.J.Lam
Gardenia anisophylla
Artocarpus elasticus Bl.
Shorea fallax Meijer
Dialium maingayi Backer
Mangifera swintonioides Kosterm.
Dead
Madhuca magnifica H.J.Lam
Sarcotheca diversifolia (Miq.) Hall.f.
Mangifera swintonioides Kosterm.
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea pinanga Scheff.
Shorea beccariana Burck.
Vatica albiramis Slooten
Diospyros sp.3
Dillenia grandifolia Wall. ex Hk.f.
Koompassia malaccensis Maing ex Benth.
Gluta wallichii (Hk.f.) Ding Hou
20 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
Species
Syzygium confertum (Korth.) Merr. & Perry
Baccaurea cf. sumatrana M.A.
Licania splendens (Korth.) Prance
Elaeocarpus stipularis Bl.
Blumeodendron elateriospermum J.J.Sm.
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck.
Agathis borneensis Warb.
Shorea beccariana Burck.
Gymnacranthera contracta Warb.
Hydnocarpus kunstleri (King) Warb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Brakenridgea palustris Bartell.
Santiria griffithii (Hk.f.) Engl.
Sindora leiocarpa Backer ex de Wit
Shorea maxwelliana King
Shorea maxwelliana King
Madhuca magnifica H.J.Lam
100
80
60
40
20
0
0
4
5
91
3
94 96
93
218
2
90
88
92
1
89
87
7
85
97
83
238
199
273
242
40
20
11
80
12 13
14
111
113
112
107
109
192
106
108
193
245
247
246
243
244
272
117
110
121
122
23
21
17
79
24
18
20
16
76
40
68
62
27
61
28
34
60
29 30
60
59
133
132
131
180
130
136
58
171
31
57
142
141
167
172
174
134
135
138
139
137
176 175
177
258
257
253
250
251
252
254 255
248
60
249
128
127
67 65 64
63
66
118
25
26
119
120
126
125
123 124
115 114
70
73
71
69
72
77 75 74
105
81
78
102
82 15
84
98
211
210
99
10
86
212
101
200
237
240
239
232
231
20
270
179
201 198 196 190 188
185
204 194
181
187 186
182
195
207
191
184 183
209
214
205
215
219
227
225
224
223
222 220
226
229
95
0
35
56
143
80
38 39
36
55
46
53
148
147
37
47
54
50
40
45
51
49
52
157
264
160
149 150
145
162
262
146
144
166
260
261
259
80
266
44
48
100
41
42
43
159
158
267
100
100
80
60
40
20
0
Figure 6. Distribution of Trees in Plot RIL02
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Arecaceae
Araucariaceae
Aquifoliaceae
Apocynaceae
Annonaceae
Verbenaceae
Ulmaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rosaceae
Rhizophoraceae
Polygalaceae
Oxalidaceae
Oleaceae
Ochnaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
FAMILY
Anacardiaceae
Lauraceae
RIL02
A Field Guide to the Permanent Sample Plots • 21
22 • A Field Guide to the Permanent Sample Plots
Table 2. List of Trees in Plot RIL02
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Polydocarpus sp.
Syzygium grandis Wight.
Cratoxylon arborescens (Vahl.) Bl.
Knema furfuracea (Hk.f. ex Th.) Warb.
Cratoxylon arborescens (Vahl.) Bl.
Dead
Cephalomappa lepidotula Airy Shaw
Diospyros sp.2
Dead
Beilschmiedia kunstleri Gamble
Polydocarpus sp.
Syzygium incarnatum (Elmer) Merr.
Palaquium rostratum (Miq.) Burck.
Syzygium grandis Wight.
Tetramerista glabra Miq.
Shorea brunnescens Ashton
Licania splendens (Korth.) Prance
Lithocarpus cf. ewickii (Korth.) A. Kamus
Shorea pinanga Scheff.
Lithocarpus cf. ewickii (Korth.) A. Kamus
Tetramerista glabra Miq.
Dead
Shorea sp.1
Gluta wallichii (Hk.f.) Ding Hou
Shorea brunnescens Ashton
Dacryodes rostrata H.J.Lam
Shorea exelliptica Meijer
Calophyllum lowii Hook.f.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Palaquium calophyllum (T.et B.) Pierre
Dead
Unidentified
Myristica iners Bl.
Mezzettia parvifolia Becc.
Shorea macroptera Dyer
Lithocarpus bennetii (Miq.) Rehd.
A Field Guide to the Permanent Sample Plots • 23
Table 2. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Artocarpus elasticus Bl.
Castanopsis cantleyanus (King & Hk.f.) Rehd.
Lithocarpus pulcherrimum (King) Markgr.
Agathis borneensis War.
Shorea rubra Ashton
Teijsmaniodendron simplicifolium Merr.
Syzygium tawahense (Korth.) Merr. & Perry
Myristica beccarii Warb.
Calophyllum pulcherrimum Wall ex Planch
Shorea parvifolia Dyer ssp. parvifoli
Koompassia malaccensis Maing ex Benth.
Dipterocarpus stellatus Vesque
Eugessona utilis Becc.
Eugessona utilis Becc.
Ficus lepicarpa Bl.
Shorea atrinervosa Sym
Anisoptera megistocarpa Slooten
Cryptocarya crassinervia Miq.
Tetramerista glabra Miq.
Adinandra sarosanthera Miq.
Diospyros sp.2
Anisoptera megistocarpa Slooten
Gironniera subaequalis Planc.
Shorea parvifolia Dyer ssp. parvifoli
Madhuca malacensis (Clarke) H.J.Lam
Artocarpus odoratissima Blanco
Dyera costulata (Miq.) Hk.f.
Chionanthus cf. pubicalyx (Ridl.) Kiew
Mezzettia parvifolia Becc.
Dipterocarpus stellatus Vesque
Koompassia malaccensis Maing ex Benth.
Shorea maxwelliana King
Agathis borneensis Warb.
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Santiria apiculata Benn.
Quercus sp.
24 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Sarcotheca diversifolia (Miq.) Hall.f.
Aporusa sp.
Polydocarpus sp.
Syzygium sp.
Polydocarpus sp.
Dalbergia sp.1
Pternandra galeata (Cogn.) Miq.
Ilex sp.
Palaquium rostratum Burck.
Shorea parvifolia Dyer ssp. parvifolia
Macaranga triloba (Bl.) Muell. Arg.
Polydocarpus sp.
Cleistanthus bakonensis Airy Shaw
Myristica iners Bl.
Polydocarpus sp.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Hydnocarpus kunstleri (King) Warb.
Myristica beccarii Warb.
Swintonia glauca Engl.
Dipterocarpus stellatus Vesque
Myristica beccarii Warb.
Shorea pinanga Scheff.
Heritiera simplicifolia (Mast.) Kosterm.
Santiria tomentosa Bl.
Hydnocarpus kunstleri (King) Warb.
Dead
Lithocarpus sp.
Lithocarpus sp.
Shorea venulosa Wood ex Meijer
Shorea venulosa Wood ex Meijer
Santiria tomentosa Bl.
Calophyllum lowii Hook.f.
Shorea macroptera Dyer
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Koompassia malaccensis Maing ex Benth.
Gluta wallichii (Hk.f.) Ding Hou
A Field Guide to the Permanent Sample Plots • 25
Table 2. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Ternstroemia sp.
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Canarim littorale Bl. f. pubescens
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Cleisthanthus bakonensis Airy Shaw
Shorea exelliptica Meijer
Gymnacranthera contracta Warb.
Mangifera magnifica Kochumen
Myristica beccarii Warb.
Dipterocarpus stellatus Vesque
Santiria driffithii (Hk.f.) Engl.
Shorea brunnescens Ashton
Dialium maingayi Backer
Shorea pinanga Scheff.
Endiandra rubescens Miq.
Dacryodes incurvata (Engl.) H.J.Lam
Dyera costulata (Miq.) Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Shorea brunnescens Ashton
Pternandra galeata (Cogn.) Miq.
Artocarpus lanceifolius Roxb.
Palaquium rostratum (Miq.) Burck.
Xylopia altissima Boerl.
Mezzettia parvifolia Becc.
Swintonia glauca Engl.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Calophyllum lowii Hook.f.
Licania splendens (Korth.) Prance
Brakenridgea palustris Bartell
26 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Carallia brachyata (Lour.) Merr.
Dyera costulata (Miq.) Hk.f.
Swintonia glauca Engl.
Shorea parvifolia Dyer ssp. parvifolia
Gironniera subaequalis Planch.
Shorea exelliptica Meijer
Shorea parvifolia Dyer ssp. parvifolia
Shorea atrinervosa Sym
Shorea hopeifolia (Heim) Sym
Dipterocarpus stellatus Vesque
Swintonia glauca Engl.
Calophyllum lowii Hook.f.
Alseodaphne ceratoxylon Kosterm
Vatica rassak (Korth.) Blume
Calophyllum lowii Hook.f.
Swintonia glauca Engl.
Shorea rubra Ashton
Xanthophyllum sp.
Shorea rubra Ashton
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Dead
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Teijsmaniodendron simplicolium Merr.
Dacryodes laxa (Benn.) Lam
Hydnocarpus kunstleri King
Koompassia malaccensis Maing ex Benth.
Castanopsis tungurut DC.
Guioa diplopetala (Hassk.) Radik
Shorea parvifolia Dyer ssp. velutina
Koompassia malaccensis Maing ex Benth.
Myristica beccarii Warb.
Parishia maingayi Hook.f.
Lithocarpus bennettii (Miq.) Rehd.
Dillenia eximia Miq.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
A Field Guide to the Permanent Sample Plots • 27
Table 2. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dipterocarpus stellatus Vesque
Shorea amplexicaulis Ashton
Parishia maingayi Hook.f.
Barringtonia macrostachya
Dipterocarpus stellatus Vesque
Swintonia glauca Engl.
Calophyllum lowii Hook.f.
Baccaurea cf. sumatrana M.A.
Parinari oblongifolia Hk.f.
Canarium littorale Bl. f. pubescens
Dipterocarpus lowii Hk.f.
Dead
Myristica maxima Warb.
Dipterocarpus lowii Hk.f.
Dipterocarpus stellatus Vesque
Syzygium sp.1
Dead
Gluta wallichii (Hk.f.) Ding Hou
Cinnamomum javanicum Bl.
Shorea beccariana Burck.
Dead
Myristica maxima Warb.
Dead
Shorea maxwelliana King
Shorea beccariana Burck.
Swintonia glauca Engl.
Shorea beccariana Burck.
Dead
Myristica maxima Warb.
Myristica maxima Warb.
Dead
Unidentified
Cryptocarya crassinervia Miq.
Dipterocarpus stellatus Vesque
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Unidentified
Paratocarpus venenosus (Zoll. & Mor.) Becc.
28 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea exelliptica Meijer
Shorea beccariana Burck.
Shorea parvifolia Dyer
Shorea beccariana Burck.
Guioa diplopetala (Hassk.) Radlk.
Lithocarpus bennetii (Miq.) Rehd.)
Dyera costulata (Miq.) Hk.f.
Cryptocarya crassinervia Miq.
Artocarpus anisophyllus Miq.
Shorea agami Ashton
Hydnocarpus gracilis (v.Sl.) Sleum
Lithocarpus cantleyanus (KIng ex Hk.f.) Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Dillenia grandifolia Wall. ex Hk.f.
Shorea exelliptica Meijer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Gymnacranthera contracta Warb.
Palaquium rostratum Burck.
Poebe laevis Kosterm.
Dipterocarpus stellatus Vesque
Swintonia glauca Engl.
Palaquium calophyllum (T. et B.) Pierre
Guioa diplopetala (Hassk.) Radlk.
Gluta wallichii (Hk.f.) Ding Hou
Castanopsis oviformis Soepadmo
Litsea firma (Hk.f.) Ding Hou
Syzygium sp.1
Pouteria malaccensis (Clarke) Baeh
Artocarpus integer (Thunb.) Merr.
Hydnocarpus kunstleri (King) Warb.
Shorea exelliptica Meijer
Shorea brunnescens Ashton
Shorea brunnescens Ashton
Lophopetalum subobovatum King
Shorea venulosa Wood ex Meijer
Myristicaceae sp.
A Field Guide to the Permanent Sample Plots • 29
Table 2. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
Species
Shorea macroptera Dyer
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Dead
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Xanthophyllum affine Miq.
Shorea exelliptica Meijer
Shorea hopeifolia (Heim) Sym
Gomphia serrata (Gaertn.) Kans.
Unidentified
Xanthophyllum parvum Chod
Ziziphus angustifolius (Miq.) Hatasima
Dipterocarpus lowii Hk.f,
Shorea parvifolia Dyer ssp. parvifolia
Gonocaryum sp.
30 • A Field Guide to the Permanent Sample Plots
Table 3. List of Trees in Plot RIL03 (dbh≥10cm)
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea parvifolia Dyer
Koompassia malaccensis Maing ex Benth.
Madhuca magnifica H.J.Lam
Elaeocarpus obtusus Bl.
Cinnamomum javanicum Bl.
Litsea mappacea (Bl.) Boerl.
Vatica albiramis Slooten
Lophopetalum aff. subobovatum King
Madhuca magnifica H.J.Lam
Shorea rubra Ashton
Dalbergia sp.1
Dipterocarpus lowii Hk.f.
Polyosma kingiana Schltr.
Lithocarpus bennettii (Miq.) Rehd.
Calophyllum pulcherrimum Wall. ex Choisy.
Shorea brunnescens Ashton
Syzygium borneense Miq.
Polyalthia glauca Boerl.
Shorea venulosa Wood ex Meijer
Syzygium chloranthum (Duthie) Merr. & Perry
Artocarpus anisophyllus Miq.
Mezzettia parvifolia Becc.
Gluta walichii (Hk.f.) Ding Hou
Scaphium macropodum (Miq.) Beaume ex Heyne
Quercus sp.
Elaeocarpus sp.
Heritiera javanica Kost.
Calophyllum lowii Hook.f.
Baccaurea puberula Merr.
Litsea nidularis Gamble
Myristica maxima Warb.
unidentified
Dead
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Garcinia dioica Bl.
A Field Guide to the Permanent Sample Plots • 31
Table 3. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Aphanamixis grandifolia Bl.
Swintonia glauca Engl.
Eugenia sp.
Baccaurea kunstleri King
Madhuca magnifica H.J.Lam
Shorea rubra Ashton
Paratocarpus venenosa Becc.
Swintonia glauca Engl.
Cleistanthus bakonensis Airy Shaw
Memecylon myrsinoides Bl.
Shorea macroptera Dyer
Xanthophyllum flavescens Roxb.
Prunus arborea (Bl.) Kalkman
Shorea macroptera Dyer
Ixonanthes petiolaris Bl.
Calophyllum pulcherrimum Wall. ex Choisy.
Garcinia sp.
Gironniera subaequalis Planch.
Syzygium tawaense (Korth.) Merr. & Perry
Shorea rubra Ashton
Shorea macroptera Dyer
Calophyllum lowii Hook.f.
Ixonanthes petiolaris Bl.
Gironniera subaequalis Planch.
Cryptocarya crassinervia Miq.
Baccaurea kunstleri King
Dialium laurinum Backer
Oncosperma horridum Scheff.
Garcinia dioica Bl.
Dacryodes rugosa H.J.Lam
Gymnacranthera contracta Warb.
Shorea parvifolia Dyer
Memecylon myrsinoides Bl.
Alseodaphne insignis Gamble
Mallotus affinis M.A.
Unidentified
32 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Mallotus affinis M.A.
Shorea rubra Ashton
Irvingia malayana Oliv. ex Benn
Shorea rubra Ashton
Cleistanthus bakonensis Airy Shaw
Baccaurea kunstleri King
Cleistanthus bakonensis Airy Shaw
Dipterocarpus stellatus Vesque
Baccaurea sumatrana Muell. Arg.
Garcinia dioica Bl.
Madhuca erythrophylla (K. et G.) H.J.Lam
Knema lunduensis (Warb.) de Wilde
Shorea parvifolia Dyer
Pimeleodendron papaveroides J.J.Sm.
Polyalthia glauca Boerl.
Syzygium ochneocarpa Merr.
Popowia hirta Miq.
Elaeocarpus petiolatus Wall.
Koilodepas brevipes Merr.
Pimeleodendron papaveroides J.J.Sm.
Mallotus affinis M.A.
Cleistanthus bakonensis Airy Shaw
Vatica umbonata (Hook.f.) Burck.
Dialium laurinum Backer
Sindora leiocarpa Backer ex Heyne
Shorea parvifolia Dyer
Artocarpus anisophyllus Miq.
Cleistanthus bakonensis Airy Shaw
Santiria griffithii (Hk.f.) Engl.
Knema furfuracea (Hk.f. et Th.) Warb.
Shorea atrinervosa Sym.
Aglaia cordata Hiern.
Dehaasia firma Bl.
Dacryodes rugosa H.J.Lam
Dipterocarpus lowii Hk.f.
Adinandra borneensis Kob.
Horsfieldia sp.
A Field Guide to the Permanent Sample Plots • 33
Table 3. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Dead
Gironniera subaequalis Planch.
Cleistanthus bakonensis Airy Shaw
Garcinia celebica Bl.
Madhuca magnifica H.J.Lam
Prunus arborea (Bl.) Kalkman
Artocarpus anisophyllus Miq.
Baccaurea kunstleri King
Gymnacranthera contracta Warb.
Santiria laevigata Bl.
Vatica albiramis Slooten
Cleistanthus bakonensis Airy Shaw
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Eugenia grandis Wight.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Fagraea fragrans Roxb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Cleistanthus bakonensis Airy Shaw
Madhuca malaccensis (Clarke) H.J.Lam
Litsea mappacea (Bl.) Boerl.
Adinandra borneensis Kobuski
Syzygium chloranthum (Duthie) Merr. & Perry
Dacryodes rostrata (Bl.) Lam
Syzygium grandis Wight.
Santiria tomentosa Bl.
Adinandra cordifolia Ridley
Syzygium sp.
Dead
Tetramerista glabra Miq.
Gluta wallichii (Hk.f.) Ding Hou
Syzygium grandis Wight.
Gluta wallichii (Hk.f.) Ding Hou
Diospyros oblonga Wall. ex G.Don
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Gomphia serrata (Gaertn.) Kanis
Shorea rubra Ashton
34 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Cleistanthus bakonensis Airy Shaw
Prunus arborea (Bl.) Kalkman
Shorea macroptera Dyer
Litsea firma Bl.
Gluta wallichii (Hk.f.) Ding Hou
Canarium littorale Bl.
Aglaia ganggoo Miq.
Shorea parvifolia Dyer
Syzygium grandis Wight.
Myristica elliptica Hk.f. et Th.
Gluta wallichii (Hk.f.) Ding Hou
Macaranga tanarius M.A.
Irvingia malayana Oliv.
Cleistanthus bakonensis Airy Shaw
Canarium littorale Bl.
Shorea atrinervosa Sym
Shorea venulosa Wood ex Meijer
Syzygium chloranthum (Duthie) Merr. & Perry
Sarcotheca diversifolia (Miq.) Hall.f.
Dialium patens Backer
Ctenolophon parvifolius Oliv.
Sarcotheca diversifolia (Miq.) Hall.f.
Artocarpus anisophyllus Miq.
Madhuca magnifica H.J.Lam
Aporusa antennifera (Airy Shaw) Airy Shaw
Artocarpus anisophyllus Miq.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Syzygium incarnatum (Elmer) Merr. & Perry
Prunus arboreus (Bl.) Kalkman
Dead
Dead
Dialium laurinum Backer
Shorea venulosa Wood ex Meijer
Calophyllum lowii Hook.f.
Pouteria malaccensis (Clarke) Bach.
Dead
Shorea venulosa Wood ex Meijer
A Field Guide to the Permanent Sample Plots • 35
Table 3. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Vatica albiramis Slooten
Dead
Gymnacranthera contracta Warb.
Cleistanthus bakonensis Airy Shaw
Melanochyla pulvinervis (Bl.) Ding Hou
Ternstroemia aneura Miq.
Canarium sp.
Aphanamixis grandifolia Bl.
Dead
Artocarpus integer (Thunb.) Merr.
Gonystylus forbesii Gilg.
Myristica beccarii Warb.)
Scaphium macropodum (Miq.) Beaume ex Heyne
Vatica granulata Slooten
Diospyros confertiflora (Hiern.) Bakh.
Baccaurea sumatrana M.A.
Cryptocarya crassinervia
Cleistanthus bakonensis Airy Shaw
Gluta wallichii (Hk.f.) Ding Hou
Chionanthus pubicalyx (Ridl.) Kiew
Prunus arboreus (Bl.) Kalkm.
Dead
Xylopia elliptica Maingay
Herritiera simplicifolia (Mast.) Kosterm.
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Gymnacranthera contracta Warb.
Cryptocarya crassinervia Miq.
Gluta wallichii (Hk.f.) Ding Hou
Cleistanthus bakonensis Airy Shaw
Memecylon sp.
Gluta wallichii (Hk.f.) Ding Hou
Vatica albiramis Slooten
Gluta wallichii (Hk.f.) Ding Hou
Dehaasia firma Bl.
Dead
Polyalthia glauca Boerl.
36 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Lithocarpus cantleyanus (King. & Hk.f.) Rehd.
Shorea macroptera Dyer
Lithocarpus cantleyanus (King. & Hk.f.) Rehd.
Myristica beccarii Warb.
Cleistanthus bakonensis Airy Shaw
Scaphium macropodum (Miq.) Beumee ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Shorea brunnescens Ashton
Gymnacranthera contracta Warb.
Helicia fuscotomentosa Sueseng.
Dipterocarpus stellatus Vesque
Cleistanthus bakonensis Airy Shaw
Gluta wallichii (Hk.f.) Ding Hou
Madhuca erythrophylla (K. et G.) H.J.Lam
Diospyros sp.
Mallotus sp.1
Syzygium borneense Miq.
Sindora beccariana Backer ex de Wit
Swintonia glauca Engl.
Gluta wallichii (Hk.f.) Ding Hou
Aglaia sp.
Syzygium sp.
Cleistanthus bakonensis Airy Shaw
Madhuca magnifica H.J.Lam
Lithocarpus nieuwenhuisii (v. Seem) A. Camus
Syzygium albidiramea Merr.
Syzygium albidiramea Merr.
Litsea nidularis Gamble
Cleistanthus bakonensis Airy Shaw
Syzygium tawaense (Korth.) Merr.
Polyalthia glauca Boerl.
Dipterocarpus stellatus Vesque
Oncosperma horridum Scheff.
Mallotus affinis M.A.
Sindora leiocarpa Backer
Shorea parvifolia Dyer
A Field Guide to the Permanent Sample Plots • 37
Table 3. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Shorea agami Ashton
Mangifera swintonioides Kosterm
Ternstroemia aneura Miq.
Dyera costulata (Miq.) Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Shorea exelliptica Meijer
Gluta wallichii (Hk.f.) Ding Hou
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea brunnescens Ashton
Phoebe laevis Kosterm.
Santiria griffithii (Hk.f.) Engl.
Durio lanceolata Mast.
Aphanamixis grandifolia Bl.
Calophyllum pulcherrimum Wall. ex Choisy
Syzygium sp.
Cinnamomum javanicum Bl.
Pouteria malaccensis (Clarke) Bach.
Shorea macroptera Dyer
Lophopetalum subobovatum King.
Litsea firma Bl.
Gluta wallichii (Hk.f.) Ding Hou
Herritiera simplicifolia (Mast.) Kosterm.
Shorea macroptera Dyer
Diospyros confertiflora (Hiern.) Bakh.
Syzygium cf. borneense Miq.
Shorea pinanga Scheff.
Memecylon myrsinoides Blume
Lithocarpus cantleyanus (King. & Hk.f.) Rehd.
Vatica albiramis Slooten
Cleistanthus bakonensis Airy Shaw
Cleistanthus bakonensis Airy Shaw
Syzygium bankense (Hassk.) Merr & Perry
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Baccaurea sumatrana M.A.
Dacryodes rostrata (Bl.) Lam.
Shorea brunnescens Haerry)
38 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
307
308
309
310
311
312
313
314
315
316
317
318
319
320
321
322
323
324
325
326
327
328
329
330
331
332
333
Species
Gluta wallichii (Hk.f.) Ding Hou
Castanopsis motleyana King
Dysoxylum cauliflorum Hiern.
Cleistanthus bakonensis Airy Shaw
Koompassia malaccensis Maing ex Benth.
Phoebe laevis Kosterm.
Dehaasia firma Bl.
Calophyllum lowii Hook.f.
Licania splendens (Korth.) Prance
Brakenridgea palustris Bartell
Shorea parvifolia Dyer
Syzygium albidiramea Merr.
Dead
Shorea brunnescens Ashton
Gaertnera vaginans Merr.
Artocarpus integer (Thunb.) Merr.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Gymnacranthera contracta Warb.
Dehaasia firma Bl.
Dipterocarpus stellatus Vesque
Pometia pinnata Forst.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Atuna sp.
Dead
Shorea venulosa Wood ex Meijer
Cleistanthus bakonensis Airy Shaw
Horsfieldia sp.
Sindora coriacea Maing. ex Prain
Palaquium calophyllum (T. et B.) Pierre
Drypetes kikir Airy Shaw
Dacryodes rugosa H.J.Lam
Gymnacranthera contracta Warb.
Unidentified
Pouteria malaccensis (Clarke) Bach.
Parinari impressa Kosterm.
Gluta wallichii (Hk.f.) Ding Hou
Prunus arborea (Bl.) Kalkman
A Field Guide to the Permanent Sample Plots • 39
Table 3. Continued
Tree #
334
335
336
337
338
339
340
341
342
343
344
345
346
347
348
349
350
351
352
353
354
355
356
357
358
359
360
361
362
363
364
365
366
367
368
369
370
Species
Gluta wallichii (Hk.f.) Ding Hou
Cleistanthus bakonensis Airy Shaw
Knema latericia Elmer
Anisophyllea ferruginea Ding Hou
Gymnacranthera contracta Warb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Xanthophyllum flavescens Roxb.
Canarium littorale Bl.
Diospyros puncticulosa Bakh.
Xylopia malayana Hook.f.
Shorea agamii Ashton
Shorea venulosa Wood ex Meijer
Dillenia grandifolia Wall ex Hk.f.
Dacryodes rugosa H.J.Lam
Dillenia grandifolia Wall ex Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Syzygium borneense Miq.
Dacryodes rostrata H.J.Lam
Syzygium tawaense (Korth.) Merr. & Perry
Dead
Eugenia ohneocarpa Merr.
Canarium littorale Bl.
Gomphia serrata (Gaertn.) Kanis
Calophyllum pulcherrimum Wall ex Planch
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Teijsmaniodendron simplicifolium Merr.
Dead
Cleistanthus myrianthus Kurz.
Eugenia sp.
Baccaurea sumatrana M.A.
Shorea pauciflora King
Vatica albiramis Slooten
Prunus arborea (Bl.) Kalkman
Cleistanthus bakonensis Airy Shaw
Gironniera subaequalis Planch.
Cleistanthus bakonensis Airy Shaw
Scaphium macropodum (Miq.) Beumée ex Heyne
40 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
371
372
373
374
375
376
377
378
379
380
381
382
383
384
385
386
387
388
389
390
391
392
393
394
395
396
397
398
399
400
401
402
403
404
405
406
407
Species
Myristica maxima Warb.
Dead
Shorea rubra Ashton
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Palaquium ferox H.J.Lam
Cleistanthus bakonensis Airy Shaw
Hydnocarpus kunstleri (King) Warb.
Dysoxylum excelsum Bl.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Dipterocarpus lowii Hk.f.
Ventilago sp.
Myristica maxima Warb.
Dead
Polyalthia glauca Boerl.
Dacryodes rugosa H.J.Lam
Santiria tomentosa Bl.
Dead
Scaphium macropodum (Miq.) Beumée ex Heyne
Dipterocarpus stellatus Vesque
Gluta wallichii (Hk.f.) Ding Hou
Aporusa maingayi Hk.f.
Dacryodes rugosa H.J.Lam
Endiandra rubescens Miq.
Memecylon sp.
Ternstroemia aneura Miq.
Madhuca sp.
Santiria griffithii (Hk.f.) Engl.
Dead
Syzygium albidiramea Merr.
Mezzetia parvifolia Becc.
Garcinia dioica Bl.
Shorea rubra Ashton
Sindora coriacea Maing. ex Prain
A Field Guide to the Permanent Sample Plots • 41
Table 3. Continued
Tree #
408
409
410
411
412
413
414
415
416
417
418
419
420
421
422
423
424
425
426
427
428
429
430
431
432
433
434
435
436
437
438
439
440
441
442
443
444
Species
Unidentified
Canarium littorale Bl.
Baccaurea sumatrana M.A.
Gluta wallichii (Hk.f.) Ding Hou
Lophopetalum subobovatum King
Mezzetia parvifolia Becc.
Cleistanthus bakonensis Airy Shaw
Garcinia dioica Bl.
Memecylon sp.
Canarium littorale Bl.
Lophopetalum subobovatum King
Eugenia chloranthum (Duthie) Merr. & Perry
Vatica chloranthum (Dut)
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Litsea mappacea (Bl.) Boerl.
Madhuca magnifica H.J.Lam
Semecarpus sp.
Dead
Eugenia chloranthum (Duthie) Merr. & Perry
Canarium littorale Bl.
Baccaurea pendula Merr.
Melanochylla pulvinervis (Bl.) Ding Hou
Horsfieldia sp.
Phanera sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Lophopetalum aff. subobovatum King
Dead
Syzygium sp.
Aporusa antennifera (Airy Shaw) Airy Shaw
Gironniera subaequalis Planch.
Myristica maxima Warb.
Shorea rubra Ashton
Pometia pinnata Forst.
Aporusa falcifera Hook.f.
Dacryodes rugosa H.J.Lam
Vatica sp.
42 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
445
446
447
448
449
450
451
452
453
454
455
456
457
458
459
460
461
462
463
464
465
466
467
468
469
470
471
472
473
474
475
476
477
478
479
480
481
Species
Helicia fuscotomentosa Sueseng.
Baccaurea pendula Merr.
Eugenia sp.
Cleistanthus bakonensis Airy Shaw
Knema cinerea (Poir.) Wrb.
Sageraea elliptica (DC.) Hk.f. et Th.
Canarium littorale Bl.
Madhuca magnifica H.J.Lam
Cryptocaria crassinervis Miq.
Aphanamixis grandifolia Bl.
Memecylon myrsinoides Blume
Sarcotheca diversifolia (Miq.) Hall.f.
Baccaurea pubera Muell Arg.
Syzygium attenuatum (Miq.) Merr. & Perry
Calophyllum pulcherrimum Wall ex Planch.
Pimeleodendron papaveroides J.J.Sm.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Prunus arborea (Bl.) Kalkman
Myristica maxima Warb.
Lophopetalum subobovatum Miq.
Artocarpus anisophyllus Miq.
Gluta wallichii (Hk.f.) Ding Hou
Myristica maxima Warb.
Dead
Memecylon myrsinoides Bl.
Shorea atrinervosa Sym.
Chionanthus cf. pubicalyx (Ridl.) Kiew
Knema cinerea (Poir.) Warb.
Hydnocarpus gracilis (v.Sl.) Sleum
Myristica elliptica Hk.f. et Th.
Gluta wallichii (Hk.f.) Ding Hou
Sarcotheca diversifolia (Miq.) Hall.f.
Randia sp.
Shorea ochracea Sym.
Shorea parvifolia Dyer
Calophyllum lowii Hook.f.
Shorea parvifolia Dyer
A Field Guide to the Permanent Sample Plots • 43
Table 3. Continued
Tree #
482
483
484
485
486
487
488
489
490
491
492
493
494
495
496
497
498
499
500
501
502
503
504
505
506
507
508
509
510
511
512
513
514
515
516
517
518
Species
Adinandra cordifolia Ridley
Baccaurea pendula Merr.
Cleistanthus bakonensis Airy Shaw
Actinodaphne sp.
Madhuca magnifica H.J.Lam
Dead
Xanthophyllum flavescens Roxb.
Dipterocarpus lowii Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Horsfieldia polyspherulla (Hk.f. et King) Sind.
Baccaurea sumatrana M.A.
Calophyllum lowii Hook.f.
Oncosperma horridum Scheff.
Syzygium chloranthum (Duthie) Merr. & Perry
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Horsfieldia polyspherulla (Hk.f. et King) Sind.
Castanopsis motleyana King
Gluta wallichii (Hk.f.) Ding Hou
Pimeleodendron papaveroides J.J.Sm.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Oncosperma horridum Scheff.
Ficus hirta Vahl.
Macaranga triloba (Bl.) Muell Arg.
Horsfieldia glabra (Bl.) Warb.
Aporusa lucida (Miq.) Airy Shaw
Shorea pinanga Scheff.
Knema latericia Elmer
Dacryodes rubiginosa (Benn.) H.J.Lam
Ctenolophon parvifolius Oliv.
Shorea parvifolia Dyer
Shorea ochracea Sym.
Shorea parvifolia Dyer
Dead
Myristica iners Bl.
Shorea macroptera Dyer
Memecylon myrsinoides Blume
Shorea pinanga Scheff.
44 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
519
520
521
522
523
524
525
526
527
528
529
530
531
532
533
534
535
536
537
538
539
540
541
542
543
544
545
546
547
548
549
550
551
552
553
554
555
Species
Cleistanthus bakonensis Airy Shaw
Gluta wallichii (Hk.f.) Ding Hou
Syzygium chloranthum (Duthie) Merr.
Ptychophyxis bacciformis Croiz.
Shorea macroptera Dyer
Mezzetia parvifolia Becc.
Dipterocarpus lowii Hk.f.
Durio lanceolatus Mast.
Shorea venulosa Wood ex Meijer
Calophyllum lowii Hook.f.
Artocarpus integer (Thenb.) Merr.
Syzygium incarnatum (Elmer) Merr. & Perry
Aporusa antennifera (Airy Shaw) Airy Shaw
Cleistanthus bakonensis Airy Shaw
Dead
Shorea parvifolia Dyer
Pimeleodendron papaveroides J.J.Sm.
Aporusa falcifera Hook.f.
Dead
Pimeleodendron papaveroides J.J.Sm.
Syzygium ochneocarpa Merr.
Madhuca magnifica H.J.Lam
Cleistanthus bakonensis Airy Shaw
Myristica beccarii Warb.
Cryptocarya crassinervia Miq.
Dacryodes rubiginosa (Benn.) H.J.Lam
Diospyros confertiflora (Hiern.) Bakh.
Shorea venulosa Wood ex Meijer
Dehaasia firma Bl.
Baccaurea pendula Merr.
Madhuca magnifica H.J.Lam
Palaquium ferox H.J.Lam
Drypetes kikr Airy Shaw
Shorea atrinervosa Sym
Oncosperma horridum Scheff.
Castanopsis motleyana King
Koompassia excelsa (Becc.) Taub.
A Field Guide to the Permanent Sample Plots • 45
Table 3. Continued
Tree #
556
557
558
559
560
561
562
563
564
565
566
567
568
569
570
571
572
573
574
575
576
577
578
579
580
581
582
583
584
585
586
587
588
589
590
591
592
Species
Tetramerista glabra Miq.
Baccaurea puberula Merr.
Diospyros confertiflora (Hiern.) Bakh.
Elaeocarpus sp.
Oncosperma horridum Sheff.
Alstonia angustiloba Miq.
Ardisia colorata Roxb.
Mallotus penangensis M.A.
Dead
Hydnocarpus kunstleri (King) Warb.
Shorea brunnescens Ashton
Unidentified
Aporusa lucida (Miq.) Airy Shaw
Dehaasia firma Bl.
Myristica iners Bl.
Pelacalyx axillaris Korth.
Memecylon myrsinoides Bl.
Vatica albiramis Slooten
Cleistanthus bakonensis Airy Shaw
Tetramerista glabra Miq.
Santiria laevigata Bl.
Artocarpus anisophyllus Miq.
Palaquium ferox H.J.Lam
Adinandra borneensis kob
Carallia brachiata (Lour.) Merr.
Durio graveolens Becc.
Pometia pinnata Forst.
Diospyros malayana Becc.
Shorea macroptera Dyer
Mallotus penangensis M.A.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Tetramerista glabra Miq.
Elaeocarpus petiolatus Wall.
Diospyros oblonga Wall. ex G.Don
Santiria griffithii (Hk.f.) Engl.
Castanopsis motleyana King
Mezzetia parvifolia Becc.
46 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
593
594
595
596
597
598
599
600
601
602
603
604
605
606
607
608
609
610
611
612
613
614
615
616
617
618
619
620
621
622
623
624
625
626
627
628
629
Species
Santiria laevigata Bl.
Lophopetalum subobovatum Miq.
Dead
Gymnacranthera contracta Warb.
Myristica crassa King
Madhuca malaccensis (Clarke) H.J.Lam
Vatica albiramis Slooten
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Baccaurea puberula Merr.
Prunus arborea (Bl.) Kalkman
Castanopsis motleyana King
Cleistanthus bakonensis Airy Shaw
Ixora blumei Z. & M.
Madhuca magnifica H.J.Lam
Mezzetia parvifolia Becc.
Aporusa lucida (Miq.) Airy Shaw
Baccaurea pendula Merr.
Shorea exelliptica Meijer
Pimeleodendron papaveroides J.J.Sm.
Lophopetalum subobovatum Miq.
Gymnacranthera contracta Warb.
Vatica granulata Slooten
Gironniera subaequalis Planch.
Plectronia didyma (Roxb.) Kurz.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Baccaurea sumatrana M.A.
Memecylon sp.
Gluta wallichii (Hk.f.) Ding Hou
Baccaurea sumatrana M.A.
Shorea exelliptica Meijer
Pimeleodendron papaveroides J.J.Sm.
Dead
Unidentified
Myristica Iners Bl.
Canarium littorale Bl.
Myristica beccarii Warb.
A Field Guide to the Permanent Sample Plots • 47
Table 3. Continued
Tree #
630
631
632
633
634
635
636
637
638
639
640
641
642
643
644
645
646
647
648
649
650
651
652
653
654
655
656
657
658
659
660
661
662
663
664
665
666
Species
Ternstroemia aneura Miq.
Shorea venulosa Wood ex Meijer
Gluta wallichii (Hk.f.) Ding Hou
Meliosma nitida Bl.
Syzygium chloranthum (Duthie) Merr.
Ternstroemia aneura Miq.
Gymnacranthera contracta Warb.
Carallia brachiata (Lour.) Merr.
Syzygium grandis Wight.
Ixora blumei Z. & M.
Dipterocarpus stellatus Vesque
Litsea firma Bl.
Dacryodes incurvata (Engl.) Lam
Vatica albiramis Slooten
Cleistanthus bakonensis Airy Shaw
Litsea mappacea (Bl.) Boerl.
Licania splendens (Korth.) Prance
Dipterocarpus lowii Hk.f.
Dipterocarpus stellatus Vesque
Shorea agami Ashton
Gluta wallichii (Hk.f.) Ding Hou
Vatica albiramis Slooten
Palaquium rostratum Burck.
Lithocarpus nieuwenhuisii (v. Seem) A.Camus
Dipterocarpus lowii Hk.f.
Cleistanthus bakonensis Airy Shaw
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Prunus arborea (Bl.) Kalkm
Hydnocarpus kunstleri (King) Warb.
Shorea parvifolia Dyer
Aporusa falcifera Hook.f.
Myristica villosa Warb.
Shorea parvifolia Dyer
Cryptocarya crassinervia Miq.
Ptychophyxis bacciformis Croiz.
Gluta wallichii (Hk.f.) Ding Hou
48 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
667
668
669
670
671
672
673
674
675
676
677
678
679
680
681
682
683
684
685
686
687
688
689
690
691
692
693
694
695
696
697
698
699
700
701
702
703
Species
Irvingia malayana Oliv.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Irvingia malayana Oliv.
Dipterocarpus lowii Hk.f.
Shorea macroptera Dyer
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Swintonia glauca Engl.
Zyziphus angustifolius (Miq.) Hatusima
Madhuca magnifica H.J.Lam
Dipterocarpus stellatus Vesque
Alseodaphne sp.
Calophyllum lowii Hook.f.
Pholidocarpus sp. sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Unidentified
Dead
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Baccaurea sumatrana M.A.
Calophyllum lowii Hook.f.
Baccaurea sumatrana M.A.
Mangifera swintonioides Kosterm.
Diospyros puncticulosa Bakh.
Calophyllum lowii Hook.f.
Gluta wallichii (Hk.f.) Ding Hou
Shorea brunnescens Ashton
Lithocarpus bennetii (Miq.) Rehd.
Scaphium macrophodum (Miq.) Beumée ex Heyne
Myristica beccarii Warb.
Gluta wallichii (Hk.f.) Ding Hou
Palaquium rostratum (Miq.) Burck.
Cleistanthus bakonensis Airy Shaw
Meliosma nitida Bl.
A Field Guide to the Permanent Sample Plots • 49
Table 3. Continued
Tree #
704
705
706
707
708
709
710
711
712
713
714
715
716
717
718
719
720
721
722
723
724
725
726
727
728
729
730
731
732
733
734
735
736
737
738
739
740
Species
Dead
Ptychophyxis bacciformis Croiz.
Calophyllum lowii Hook.f.
Dead
Horsfieldia bracteata Hendl.
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata Bl.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Endospermum malaccense Benn. et. M.A.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Litsea firma Bl.
Dead
Parastemon urophyllus (A.DC.) A.DC.
Memecylon costatum Miq.
Dialium maingayi Backer
Anisophylla ferruginea Ding Hou
Horsfieldia grandis (Hk.f.) Warb.
Microcos sp.
Gymnacranthera contracta Warb.
Mallotus pinangensis M.A.
Anisoptera megistocarpa Slooten
Madhuca erythrophylla (K.et. G.) H.J.Lam
Madhuca magnifica H.J.Lam
Dacryodes incurvata (Engl.) H.J.Lam
Shorea parvifolia Dyer
Myristica iners Bl.
Dead
Memecylon myrsinoides Blume
Dacryodes laxa (Benn.) H.J.Lam
Shorea parvifolia Dyer
Hydnocarpus gracilis (v. Sl.) Sleum
Prunus arboreus (Bl.) Kalkman
Artocarpus anysophyllus Miq.
Pometia pinnata Forst.
Alseodaphne ceratoxylon Kosterm
50 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
741
742
743
744
745
746
747
748
749
750
751
752
753
754
755
756
757
758
759
760
761
762
763
764
765
766
767
768
769
770
771
772
773
774
775
776
777
Species
Ilex sp.
Shorea parvifolia Dyer
Gluta wallichii (Hk.f.) Ding Hou
Macaranga gigantea M.A.
Shorea parvifolia Dyer
Artocarpus anysophyllus Miq.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Pimeleodendron papaveroides J.J.Sm.
Santiria tomentosa Bl.
Dead
Shorea parvifolia Dyer
Scaphium macrophodum (Miq.) Beumée ex Heyne
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Ilex sp.
Shorea macroptera Dyer
Elaeocarpus polystachyus Wall
Dead
Elaeocarpus polystachyus Wall
Shorea brunnescens Ashton
Dead
Shorea parvifolia Dyer
Syzygium albidiramea Merr.
Cleistanthus bakonensis Airy Shaw
Dead
Aporusa falcifera Hook.f.
Cleistanthus bakonensis Airy Shaw
Scaphium macropodum (Miq.) Beumée ex Heyne
Palaquium calophyllum Pierre
Dyera costulata (Miq.) Hk.f.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea parvifolia Dyer
Guioa diplopetala (Hassk.) Ridl.
Litsea nidularis Gamble
Calophyllum lowii Hook.f.
Shorea maxwelliana King
Syzygium sp.
Garcinia dioica Bl.
A Field Guide to the Permanent Sample Plots • 51
Table 3. Continued
Tree #
778
779
780
781
782
783
784
785
786
787
788
789
790
791
792
793
794
795
796
797
798
799
800
Species
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Cleistanthus bakonensis Airy Shaw
Sarcotheca diversifolia (Miq.) Hall. f.
Horsfieldia polyspherulla (Hk.f. et King) Sincl.
Calophyllum lowii Hook.f.
Shorea pauciflora King.
Podocarpus neriifolius D.Don
Cleistanthus bakonensis Airy Shaw
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Heritiera javanica Kost.
Memecylon costatum Miq.
Gluta cf. wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Shorea rubra Ashton
Shorea parvifolia Dyer
Zizyphus angustifolius (Miq.) Hatusima
Mangifera swintonioides Kosterm.
Guioa diplopetala (Hassk.) Ridl.
Prunus arborea (Bl.) Kalkman
Shorea pauciflora Dyer
Dillenia grandifolia Wall ex HK.f.
Guioa diplopetala (Hassk.) Ridl.
100
80
60
40
20
0
0
0
1
2
74 75
165
166
167
169
172
171
163
78
63
71 68
69
70
72
73
168
164
170
76
64
67
175
20
3
20
62
82
58
57
181
182
184
55
80
4
159
160
158
180
81
161
179
187
186
56
66 65
79
83
162
178
177
176
174
5
7
91
90
89
6
92
98
154
85
60
93
88
87
40
61
86
150
155
188
189
152 151
153
156
185
183
40
103
148
190
95
94
99
12
10
13
200
147
145
144
141
53
52
49
111
15
60
14
48
113 112
223
47
45
16
17
43
114
21
19
131
129
80
20
132
136
133
18
22
42
46
80
40
23
222
38
31
28
123
100
26
33
37
36
30
32
29
100
34
35
122
121
24 25
39
120
125
126 124
128
221
118
210
217 216
220
218
208
219
215
209
213
214
211
207
115
116
134
135
206
140 139
138
201
205
202
203
204
106 107
108
143
104 105
51
50
96
98
60
196
146
197
195
191
101 100 102
149
192
193 194
100
80
60
40
20
0
Figure 7. Distribution of Trees in Plot RIL04
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Erythroxylaceae
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Annonaceae
Ulmaceae
Tiliaceae
Thymelaeaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rutaceae
Rosaceae
Rhizophoraceae
Rhamnaceae
Polygalaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
FAMILY
Anacardiaceae
Lauraceae
RIL04
52 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 53
Table 4. List of Trees in Plot RIL04
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Xanthophyllum amoenum Chod.
Shorea ovalis (Korth.) Bl.
Hydnocarpus kunstleri (King) Warb.
Artocarpus kemando Miq.
Neesia synandra Mast.
Shorea fallax Meijer
Macaranga hosei King ex Hk.f.
Shorea macroptera Dyer
Shorea fallax Meijer
Myristica maxima Warb.
Unidentified
Shorea pauciflora Dyer
Dialium maingayi Backer
Atuna sp.
Pholidocarpus sp.
Dacryodes rostrata (Bl.) Lam
Shorea parvifolia Dyer
Madhuca kingiana (Clarke) H.J.Lam
Agathis borneensis Warb.
Madhuca kingiana (Clarke) H.J.Lam
Shorea brunnescens Ashton
Dipterocarpus lowii Hk.f.
Madhuca kingiana (Clarke) H.J.Lam
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea parvifolia Dyer
Vatica albiramis Slooten
Dead
Artocarpus integer (Thunb.) Merr.
Hydnocarpus kunstleri (King) Warb.
Shorea parvifolia Dyer
Mangifera swintonioides Kosterm.
Shorea exelliptica Meijer
Mezzettia parviflora Becc.
Shorea brunnescens Ashton
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Mangifera magnifica Kochumen
54 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Adinandra sarosanthera Miq.
Archidendron clipearia (Jack.) Nielsen
Pholidocarpus sp.
Dead
Palaquium sp.1
Xylopia altisima Boerl.
Dead
Koompassia malaccensis Maing. ex Benth.
Aglaia ganggo Miq.
Shorea macroptera Dyer
Tetramerista glabra Miq.
Gymnacranthera contracta Warb.
Aporusa lucida (Miq.) Airy Shaw
Gironniera subaequalis Planch.
Dacryodes incurvata (Engl.) Lam
Pternandra galeata Cogn.
Unidentified
Trigonopleura malayana Hook.f.
Elaeocarpus valetani Kochr.
Koompassia malaccensis Maing ex Benth.
Gymnacranthera contracta Warb.
Unidentified
Shorea macroptera Dyer
Dacryodes incurvata (Engl.) Lam
Shorea macroptera Dyer
Artocarpus kemando Miq.
Carallia brachyata (Lour) Merr.
Shorea fallax Meijer
Shorea brunnescens Ashton
Celtis philipensis Blanco
Shorea pinanga Scheff.
Syzygium incarnatum (Elmer) Merr & Perry
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Madhuca kingiana (Clarke) H.J.Lam
Koompassia malaccensis Maing ex Benth.
Xanthophyllum amoenum Chod.
A Field Guide to the Permanent Sample Plots • 55
Table 4. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Tetramerista glabra Miq.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea pinanga Scheff.
Syzygium chloranthum (Duthie) Bakh.
Dacryodes incurvata (Engl.) H.J.Lam
Parashorea smythiesii Wyatt-Smith & Ashton
Knema latericia Elmer
Scaphium macropodum (Miq.) Beumee
Dead
Xathophyllum rufum Benth.
Litsea machilifolia Gamble
Tetramerista glabra Miq.
Ctenolophon parvifolius Oliv.
Chionanthus pubicalyx (Ridl.) Kiew
Baccaurea sumatrana M.A.
Gluta wallichii (Hk.f.) Ding Hou
Pouteria malaccensis (Clarke) Baeh
Litsea nidularis Gamble
Castanopsis motleyana King
Myristica beccarii Warb.
Madhuca magnifica H.J.Lam
Dead
Shorea pinanga Scheff.
Baccaurea sumatrana M.A.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea brunnescens Ashton
Dipterocarpus lowii Hk.f.
Shorea brunnescens Ashton
Shorea multiflora (Burck.) Sym
Shorea parvifolia Dyer
Celtis philipensis Blanco
Canarium littorale Bl.
Dipterocarpus lowii Hk.f.
Dead
Dead
Shorea exelliptica Meijer
56 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Syzygium bankense (Hassk.) Merr. & Perry.
Dialium patens Backer
Sindora leiocarpa Backer ex de Wit
Dipterocarpus lowii Hk.f.
Mangifera magnifica Kochumen
Dead
Agathis borneensis Warb.
Myristica iners Bl.
Mangifera magnifica Kochumen
Artocarpus odoratisimus Blanco
Shorea brunnescens Ashton
Gymnacranthera contracta Warb.
Lithocarpus ulceolaris (Jack) Merr.
Herritiera javanica (Bl.) Kosterm
Microcos cinnamomiifolia Burr.
Shorea brunnescens Ashton
Pimeleodendron papaveroides J.J.Sm.
Parastemon urophyllus (A.DC.) A.DC.
Dead
Aquilaria malaccensis Lamk.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Carallia brachiata (Lour.) Merr.
Tetrameristra glabra Miq.
Parastemon urophyllus (A.DC.) A.DC.
Ternstroemia aneura Miq.
Dead
Ternstroemia aneura Miq.
Dillenia grandifolia Wall ex Hk.f.
Palaquium rostratum Burck.
Shorea venulosa Wood ex Meijer
Dead
Dialium patens Backer
Vatica vinosa Ashton
Shorea macroptera Dyer
Tetramerista glabra Miq.
Oncosperma horridum Sheff.
Shorea macroptera Dyer
A Field Guide to the Permanent Sample Plots • 57
Table 4. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Syzygium attenuatum (Miq.) Merr. & Perry
Sterculia cordata Bl.
Pouteria malaccensis (Clarke) Baehni
Cratoxylum arborescens
Litsea grandis (Wall. ex Nees) Hook.f.
Myristica maxima Warb.
Xylopia elliptica Maingay
Gluta wallichii (Hk.f.) Ding Hou
Dead
Anisoptera megistocarpa Slooten
Heritiera simplicifolia (Mast.) Kosterm.
Vatica rassak (Korth.) Blume
Shorea rubra Ashton
Shorea parvifolia Dyer
Pometia pinnata Forst.
Baccaurea sumatrana Muell Arg.
Shorea pinanga Scheff.
Eugenia incarnatum (Elmer) Merr. & Perry
Blumeodendron elateriospermum J.J.Sm.
Oncosperma horridum Scheff.
Licania splendens (Korth.) Prance
Sindora beccariana Backer ex de Wit
Licania splendens (Korth.) Prance
Shorea atrinervosa Sym
Dead
Shorea parvifolia Dyer
Durio dulcis Becc.
Ziziphus angustifolius (Miq.) Hatusima
Moultonianthus leembrugianus (Boerl. & Koord.) Steenis
Shorea macroptera Dyer
Dacryodes rubiginosa (Benn.) H.J.Lam
Oncosperma horridum Scheff.
Syzygium grandis Wight.
Knema latericia Elmer
Shorea macroptera Dyer
Hydnocarpus kunstleri (King) Warb.
Shorea beccariana Burck.
58 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Diospyros sp.
Koompassia malaccensis Maing ex Benth.
Palaquium ferox H.J.Lam
Madhuca malaccensis (Clarke) H.J.Lam
Clophyllum lowii Hook.f.
Ternstroemia aneura Miq.
Drypetes kikir Airy Shaw
Tetractomia obovata
Shorea macroptera Dyer
Artocarpus dadah Miq.
Mangifera swintonioides Kosterm
Pouteria malaccensis (Clarke) Baehni
Dead
Dead
Pouteria malaccensis (Clarke) Baehni
Myristica villosa Warb.
Shorea fallax Meijer
Dipterocarpus confertus Sloot.
Erythroxylon cuneatum (Miq.) Kurz.
Polydocarpus sp.
Tetramerista glabra Miq.
Madhuca magnifica H.J.Lam
Artocarpus anisophyllus Miq.
Madhuca magnifica H.J.Lam
Dialium modestum (v.Steenis) Stegaert.
Oncosperma horridum Scheff.
Dead
Vatica vinosa Ashton
Tetramerista glabra Miq.
Ternstroemia aneura Miq.
Gluta wallichii (Hk.f.) Ding Hou
Ternstroemia magnifica Staff.
Litsea nidularis Gamble
Gymnacranthera contracta Warb.
Madhuca magnifica H.J.Lam
Shorea exelliptica Meijer
Xanthophyllum rufum A.W. Bennett
A Field Guide to the Permanent Sample Plots • 59
Table 4. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
Species
Artocarpus anisophyllus Meijer
Shorea rubra Ashton
Anisoptera megistocarpa Slooten
Lithocarpus lucidus (Roxb.) Rehd.
Shorea parvifolia Dyer
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Calophyllum lowii Hook.f.
Gluta wallichii (Hk.f.) Ding Hou
Magnolia gigantifolia (Miq.) Noot
Xylopia caudata Hk.f ex Th.
Artocarpus anisophyllus Miq.
Madhuca kingiana (Clarke) H.J.Lam
Artocarpus integer (Thunb.) Merr.
Gluta wallichii (Hk.f.) Ding Hou
Shorea pinanga Scheff.
Myristica crassa King
Dead
Dehaasia firma Bl.
Canarium littorale Bl.
Diospyros confertiflora (Hiern.) Bakh.
Pimeleodendron papaveroides J.J.S.
Shorea parvifolia Dyer
Dead
Unidentified
100
80
60
40
20
0
0
0
265
151
150
153
294
258
264
52
155
67
20
208
100
165
164
204
58
230
161
259
198 154 156
302
28
249
162
94
163
96 199
20
272
277
250
191
298
237
296
157
40
213 290
261
72
236
235
288
182
246
89
301
276
158
159
278
40
113
103
147
93
257
66
216
284
203
211
73
180
144
145
45
283
60
196
268
54
121
300
75
62
273
183
60
287
141
63
263
91
285
128
197
140
220
271
202
229
124
251
166
80
80
185
281
219
270
269
214
291
42 225
205 274
46
279 184 280
100
207
252
215
100
100
80
60
40
20
0
Figure 8. Distribution of Tree in Plot RIL05
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Clusiaceae
Burseraceae
Arecaceae
Annonaceae
Verbenaceae
Theaceae
Sterculiaceae
Simarubaceae
Sapotaceae
Rosaceae
Podocarpaceae
Olacaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
FAMILY
Anacardiaceae
Lauraceae
RIL05
60 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 61
Table 5. List of Trees in Plot RIL05
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Agathis borneensis Warb.
Shorea atrinervosa Sym
Santiria laevigata Bl.
Memecylon myrsinoides Blume
Aporusa maingayi
Gluta wallichii (Hk.f.) Ding Hou
Myristica maxima Warb.
Garcinia rostrata T. ex B.
Tristaniopsis whiteana (Griff.) Wilson
Gluta wallichii (Hk.f.) Ding Hou
Tristaniopsis whiteana (Griff.) Wilson
Baccaurea sumatrana Muell Arg.
Santiria laevigata Bl.
Dead
Santiria laevigata Bl.
Mezzetia havilandii (Boerl.) Ridl.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Quercus gmelliflora Bl.
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes incurvata (Engl.) H.J.Lam
Shorea atrinervosa Sym
Castanopsis motleyana King
Litsea mappacea (Bl.) Boerl.
Shorea macroptera Dyer
Xanthophyllum parvum Chod.
Mezzettia parvifolia Becc.
Dacryodes rostrata (Bl.) Lam
Tetramerista glabra Miq.
Alseodaphne ceratoxylon Kosterm
Myristica villosa Warb.
Barringtonia macrostachya (Jack.) Kurz.
Lithocarpus bennettii (Miq.) Rehd.
Baccaurea puberula Merr.
Ilex sp.
Neesia synandra Mast.
Diospyros sp.1
Dead
62 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Gluta wallichii (Hk.f.) Ding Hou
Neesia synandra Mast.
Teijsmaniodendron simplicifolium Merr.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea venulosa Wood ex Meijer
Madhuca malaccensis (Clarke) H.J.Lam
Cleistanthus bakonensis Airy Shaw
Syzygium chloranthum (Duthie) Merr & Perry
Shorea venulosa Wood ex Meijer
Cleistanthus bakonensis Airy Shaw
Dead
Baccaurea sumatrana Muell Arg.
Aglaia ganggo Miq.
Litsea nidularis Gamble
Teijsmaniodendron simplicifolium Merr.
Myristica maxima Warb.
Podocarpus neriifolius D.Don
Mezzettia parvifolia Becc.
Macaranga motleyana (M.A.) M.A.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Palaquium cochlearifolium v. Royen
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Palaquium cochlearifolium v. Royen
Gymnacranthera contracta Warb.
Teijsmaniodendron simplicifolium Merr.
Scaphium macropodum (Miq.) Beumée ex Heyne
Cleistanthus bakonensis Airy Shaw
Unidentified
Syzygium caudatilimbum (Merr.) Merr.
Dipterocarpus lowii Hk.f.
Harmandia mekongensis Pierre
Prunus arborea (Bl.) Kalkman
Diospyros puncticulosa Bakh.
Cleistanthus sp.
Shorea venulosa Wood ex Meijer
Palaquium cochlearifolium van Royen
Mallotus griffithianus (Muell. Arg.) Hook.f.
A Field Guide to the Permanent Sample Plots • 63
Table 5. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Dialium maingayi Backer
Ternstroemia aneura Miq.
Shorea maxwelliana King
Agathis borneensis War.
Shorea rubra Ashton
Lophopetalum subobovatum King
Shorea maxwelliana King
Aglaia ganggo Miq.
Sindora coriacea Maing. ex Prain
Dacryodes rugosa (Bl.) Lam
Baccaurea sumatrana Muell. Arg.
Polyosma kingiana Schtr.
Vatica umbonata (Hook.f.) Burck.
Shorea rubra Ashton
Shorea maxwelliana King
Syzygium ochneocarpa Merr.
Scaphium macropodum (Miq.) Beumée ex Heyne
Madhuca malaccensis (Clarke) H.J.Lam
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Shorea atrinervosa Ashton
Gymnacranthera contracta Warb.
Cleistanthus bakonensis Airy Shaw
Endiandra sp.
Shorea macroptera Dyer
Actinodaphne borneensis Meissn.
Memecylon myrsinoides Blume
Baccaurea pendula Merr.
Myristica maxima Warb.
Baccaurea sumatrana M.A.
Shorea parvifolia Dyer
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Endiandra rubescens Miq.
Dead
Diospyros confertiflora (Hiern.) Bakh.
Cleistanthus bakonensis Airy Shaw
Shorea atrinervosa Sym
64 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Memecylon sp.
Scaphium macropodum (Miq.) Beumée ex Heyne
Myristica maxima Warb.
Dead
Syzygium caudatilimbum (Merr.) Merr.
Canarium littorale Bl.
Xanthophyllum flavrescens Roxb.
Shorea beccariana Burck.
Dead
Cleistanthus bakonensis Airy Saw
Gluta wallichii (Hk.f.) Ding Hou
Dead
Buchanania arborescens (Bl.) Bl.
Mezzettia parvifolia Becc.
Madhuca malaccensis (Clarke) H.J.Lam
Dead
Cleistanthus bakonensis Airy Shaw
Dead
Eugenia sp.
Barringtonia macrostachya (Jack.) Kurz.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Ternstroemia aneura Miq.
Madhuca malaccensis (Clarke) H.J.Lam
Atuna racemosa Rafin
Cleistanthus sp.
Syzygium caudatilimbum (Merr.) Merr.
Gymnacranthera contracta Warb.
Mezzettia parvifolia Becc.
Shorea maxwelliana King
Shorea artivervosa Sym
Shorea rubra Ashton
Artocarpus integer (Thunb.) Merr.
Shorea maxwelliana King
Parinari obolongifolia Hk.f.
Calophyllum lowii Hook.f.
A Field Guide to the Permanent Sample Plots • 65
Table 5. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea maxwelliana King
Pouteria malaccensis (Clarke) Bach.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Dead
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea pinanga Scheff.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Artocarpus integer (Thunb.) Merr.
Myristica beccarii Warb.
Harmandia mekongensis Pierre
Diospyros evena Bakh.
Shorea pinanga Scheff.
Calophyllum pulcherrimum Wall. ex Choisy et Triana
Shorea maxwelliana King
Shorea brunnescens Ashton
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Shorea brunnescens Ashton
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea exelliptica Meijer
Harmandia mekongensis Pierre
Parinari oblongifolia Hk.f.
Sindora beccariana Backer ex de Wit
Shorea rubra Ashton
Agathis borneensis Warb.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea exelliptica Meijer
Aporusa antennifera (Airy Shaw) Airy Shaw
Palaquium cochlearifolium van Royen
Dipterocarpus lowii Hk.f.
Madhuca erythrophylla (K. et G.) H.J.Lam
Semecarpus rufovelutinus Ridley
Hydnocarpus sp.
Dacryodes rubiginosa (Benn.) H.J.Lam
Artocarpus integer (Thunb.) Merr.
66 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Agathis borneensis Warb.
Santiria tomentosa Bl.
Palaquium rostratum Burck.
Agathis borneensis Warb.
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Diospyros evena Bakh.
Palaquium cochlearifolium van Royen
Syzygium bankense (Hassk.) Merr. & Perry
Myristica beccarii Warb.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Myristica beccarii Warb.
Palaquium rostratum (Miq.) Burck.
Memecylon myrsinoides Blume
Irvingia malayana Oliv ex Benn
Shorea beccariana Burck.
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Xylopia altissima Boerl.
Oncosperma horridum Scheff.
Baccaurea sumatrana Muell. Arg.
Diospyros evena Bakh.
Syzygium bankense (Hassk.) Merr. & Perry
Palaquium cochlearifolium van Royen
Shorea rubra Ashton
Shorea maxwelliana King
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Syzygium chloranthum (Duthie) Merr. & Perry
Palaquium cochlearifolium van Royen
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Syzygium sp.
Blumeodendron elateriospermum J.J.S.
Myristica beccarii Warb.
A Field Guide to the Permanent Sample Plots • 67
Table 5. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea parvifolia Dyer
Harmandia makongensis Pierre
Syzygium ochneocarpa Merr.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Unidentified
Ternstroemia aneura Miq.
Baccaurea sumatrana M.A.
Dead
Elaeocarpus sp.
Calophyllum pulcherrimum Wall ex Planch.
Shorea exelliptica Meijer
Garcinia dioica Bl.
Koompassia malaccensis Maing ex Benth.
Prunus arborea (Bl.) Kalkm.
Dead
Agathis borneensis Warb.
Shorea exelliptica Meijer
Calophyllum pulcherrimum Wall. ex Choisy et Triana
Syzygium caudatilimbum (Merr.) Merr. & Perry
Dead
Dead
Tristania whiteana (Griff.) Wilson & Waterhouse
Palaquium rostratum (Miq.) Burck.
Shorea venulosa Wood ex Meijer
Elaeocarpus cf. petiolatus Wall.
Diospyros evena Bakh.
Syzygium sp.
Shorea venulosa Wood ex Meijer
Palaquium rostratum (Miq.) Burck.
Diospyros evena Bakh.
Palaquium sp.
Mezzettia parvifolia Becc.
Neolitsea cassiaefolia (Bl.) Merr.
Palaquium rostratum (Miq.) Burck.
Tristaniopsis whiteana (Griff.) Wilson & Waterhouse
Syzygium chloranthum (Duthie) Merr. & Perry
68 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Diospyros evena Bakh.
Diospyros evena Bakh.
Syzygium sp.
Tetramerista glabra Miq.
Shorea macroptera Dyer
Shorea macroptera Dyer
Unidentified
Irvingia malayana Oliv ex Benn.
Gymnacranthera contracta Warb.
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Shorea maxwelliana King
Shorea maxwelliana King
Canarium littorale Bl.
Shorea maxwelliana King
Dipterocarpus lowii Hk.f.
Koompassia malaccensis Maing ex Benth.
Shorea parvifolia Dyer
Myristica beccarii Warb.
Harmandia mekongensis Pierre
Shorea brunnescens Ashton
Scaphium macropodum (Miq.) Beumee ex Heyne
Memecylon mysinoides Blume
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata H.J.Lam
Pternandra caerulescens Jack.
Dead
Quercus gmelliflora Bl.
Santiria oblongifolia Bl.
Dialium patens Backer
Syzygium chloranthum (Duthie) Merr. & Perry
Shorea venulosa Wood ex Meijer
Dead
Dead
Blumeodendron elateriospermum J.J.S.
Ventilago sp.
Myristica elliptica Hk.f. et Th.
A Field Guide to the Permanent Sample Plots • 69
Table 5. Continued
Tree #
297
298
299
300
301
302
303
304
305
Species
Unidentified
Syzygium incarnatum (Elmer) Merr. & Perry
Dead
Shorea parvifolia Dyer
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea maxwelliana King
Madhuca erythrophylla (K. et G.) H.J.Lam
Gnetum cuspidatum Bl.
Shorea venulosa Wood ex Meijer
100
80
60
40
20
0
0
0
122
7
53
44
108
86
208
193
66
120
107
87
164
95
116
163
97
85
96
74
248
33
20
45
118
48
25
153
229
20
8
35
20
22
183
81
41
40
12
11
182
6
174
160
154
169
125 142
172
180
36
17
133
121
136
244
224
39
68
156
168
40
54
4
5
243
60
239
101
50
51
52
246
60
149
46
38
59
148
200
40
185
166
91 92
80
80
129
100
155
171 159
131 150
141
94
14
139
9
100
34
3
231
43
75
100
13
235
78
93
238
15
225
161
223
247
88
175
10
57 167
242
73
79
126
152
98
124
184
151
67
21
19
82
100
80
60
40
20
0
Figure 9. Distribution of Trees Plot RIL06
Dilleniaceae
Crypteroniaceae
Burseraceae
Bombacaceae
Arecaceae
Annonaceae
Myrtaceae
Myristicaceae
Meliaceae
Lauraceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
FAMILY
Anacardiaceae
Dipterocarpaceae
RIL06
70 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 71
Table 6. List of Trees in Plot RIL06
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Madhuca malaccensis (Clarke) H.J.Lam.
Madhuca malaccensis (Clarke) H.J.Lam.
Pimeleodendron papaveroides J.J.S.
Shorea macroptera Dyer
Shorea exelliptica Meijer
Vatica maingayi Dyer
Gluta wallichii (Hk.f.) Ding Hou
Syzygium grandis Wight.
Santiria griffithii (Hk.f.) Engl.
Knema latericia Elmer
Vatica maingayi Dyer
Dipterocarpus stellatus Vesque
Santiria laevigata Bl.
Syzygium sp.
Syzygium grandis Wight.
Dead
Vatica maingayi Dyer
Tetramerista glabra Miq.
Syzygium chloranthum (Duthie) M. & P.
Shorea fallax Meijer
Lithocarpus ewyckii (Korth.) Rehd.
Syzygium incarnatum (Elmer) M. & P.
Dead
Heritiera javanica (Bl.) Kosterm.
Vatica vinosa Ashton
Ochanostachys amentacea Mast.
Polyalthia lateriflora (Bl.) King
Tetramerista glabra Miq.
Irvingia malayana Oliv ex Benn.
Lithocarpus pseudokunstleri A. Camus
Nephelium uncinatum Leenh.
Xerospermum noronhianum Bl.
Alseodaphne insignis Gamble
Cleistanthus bakonensis Airy Shaw
Vatica vinosa Ashton
Vatica vinosa Ashton
Xanthophyllum amoenum Chod.
72 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Castanopsis motleyana King
Crypteronia griffithii Clarke
Lithocarpus conocarpus (Oudem.) Rehd.
Shorea macroptera Dyer.
Microcos cinnamomifolia Burr.
Drypetes kikir Airy Shaw
Mangifera magnifica Kochumen
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus ewyckii (Korth.) Rehd.
Helicia fuscotomentosa Susseng.
Shorea exelliptica Meijer
Syzygium ochneocarpa Merr.
Lithocarpus ewyckii (Korth.) Rehd.
Lithocarpus ewyckii (Korth.) Rehd.
Artocarpus integer (Thunb.) Merr.
Shorea exelliptica Meijer
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Nephelium lappaceum Miq.
Teijsmaniodendron simplcifolium Merr.
Myristica beccarii Warb.
Syzygium chloranthum (Duthie) M. & P.
Dipterocarpus stellatus Vesque
Baccaurea puberula Merr.
Xanthophyllum amoenum Chod.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Dead
Vatica vinosa Ashton
Teijsmaniodendron simplicifolium Merr.
Vatica vinosa Ashton
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Phoebe laevis Kostrm.
Pentace laxiflora Merr.
Shorea parvifolia Dyer
Tetramerista glabra Miq.
Gironniera subaequalis Planch.
Trigonopleura malayana Hk.f.
Shorea parvifolia Dyer
A Field Guide to the Permanent Sample Plots • 73
Table 6. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Santiria laevigata Bl.
Dipterocarpus stellatus Vesque
Dacryodes incurvatum. H.J.Lam
Diospyros laevigata Bakh.
Ptychophyxis bacciformis Croiz.
Teijsmaniodendron simplicifolium Merr.
Castanopsis tungurut (Bl.) A.Dc.
Artocarpus anisophyllus Miq.
Heritiera simplicifolia (Mast.) Kosterm
Dead
Dipterocarpus stellatus Vesque
Mangifera swintonioides Kosterm.
Vatica umbonata (Hook.f.) Burck
Drypetes kikir Airy Shaw
Teijsmaniodendron simplicifolium Merr.
Teijsmaniodendron simplicifolium Merr.
Artocarpus odoratissima Blanco
Artocarpus anisophyllus Miq.
Santiria laevigata Bl.
Dillenia grandifolia Wall. ex Hk.f
Shorea parvifolia Dyer .
Shorea beccariana Burck
Shorea parvifolia Dyer
Dialium maingayi Baker
Madhuca malaccensis (Clarke) H.J.Lam
Santiria laevigata Bl.
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Adinandra borneensis Kob.
Vatica vinosa Ashton
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Shorea rubra Ashton
Gluta wallichii (Hk.f) Ding Hou
Dead
Mangifera swintonioides Kosterm.
Hydnocarpus kunstleri (King & Hk.f) Rehd.
74 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Agathis borneensis Warb.
Gluta wallichii (Hk.f.) Ding Hou
Licania splendens (Korth) Prance
Shorea macroptera Dyer
Mangifera swintonioides Konsterm.
Durio lanceolata Mast.
Shorea rubra Ashton
Gomphia serrata (Gaertn.) Kanis
Shorea parvifolia Dyer
Mezzetia parviflora Becc.
Swintonia glauca Engl.
Shorea macroptera Dyer
Baccaurea sumatrana M.A.
Castanopsis motleyana King
Artocarpus integer (Thunb.) Merr.
Myrisytica iners Bl.
Madhuca magnifica H.J.Lam
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Teijsmaniodendron simplicifolium Merr.
Shorea parvifolia Dyer
Chionanthus ellipticus Bl.
Tetramerista glabra Miq.
Cyathocalyx sp.
Shorea macroptera Dyer
Vatica vinosa Ashton
Durio testudinarum Becc.
Pellacalyx sp.
Crypteronia griffithii Clarke
Castanopsis motleyana King
Shorea exelliptica Meijer
Shorea macroptera Dyer
Prunus arborea (Bl.) Kalkm.
Vatica vinosa Ashton
Dyera eximia Val.
Crypteronia griffithii Clarke
A Field Guide to the Permanent Sample Plots • 75
Table 6. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Crypteronia griffithii Clarke
Oncosperma horridum Scheff.
Macaranga conifera (Zoll.) M. A.
Dacryodes incurvata (Engl.) H.J.Lam
Shorea agami Ashton
Shorea macroptera Dyer
Santiria larvigata Bl.
Litsea firma Bl.
Palaquium cochlearifolium van Royen
Helicia fuscotomentosa Sueseng.
Hydnocarpus anomala (Merr.) Sleum.
Shorea fallax Meijer
Oncosperma horridum Scheff.
Dead
Mangifera swintonioides Kosterm.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Koompassia malaccensis Maing ex Benth.
Gymnacranthera contracta Warb.
Dehaasia firma Bl.
Shorea macroptera Dyer
Syzygium sp.
Santiria laevigata Bl.
Lithocarpus blumeanus (Korth.) Rehd.
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea ovalis (Korth.) Bl.
Myristica iners Bl.
Madhuca magnifica H.J.Lam
Dead
Palaquium ferox H.J.Lam
Prunus arborea (Bl.) Kalkm.
Mezzetia parviflora Becc.
Tetramerista glabra Miq.
Aglaia cordata Hiern.
Alseodaphne insignis Gamble
Pimeleodendron papaveroides J.J.S.
Koompassia malaccensis Maing ex Benth.
76 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Madhuca magnifica H.J.Lam
Dipterocarpus lowii Hook.f.
Dead
Shorea parvifolia Dyer ssp. parvifolia
Oncosperma horridum Scheff.
Artocarpus odoratissima Blanco
Oncosperma horridum Scheff.
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes incurvata (Engl.) H.J.Lam.
Shorea maxwelliana King
Scaphium macropodum (Miq.) Beumee
Shorea maxwelliana King
Mezzetia parviflora Becc.
Agathis borneensis Warb.
Vatica maingayi Dyer
Agathis borneensis Warb.
Syzygium bankesis (Hassk) Merr. & Perry
Dead
Madhuca erytrophylla (K.ex G.) H.J.Lam
Shorea beccariana Burck.
Mezzetia parviflora Becc.
Dacryodes incurvata (Engl.) H.J.Lam.
Swintonia glauca Engl.
Syzygium chloranthum (Duthie) M. & P.
Shorea parvifolia Dyer
Santiria griffithii (Hk.f.) Engl.
Licania splendens (Korth.) Prance
Dialium maingayi Baker
Vatica umbonata (Hook. f.) Burck
Swintonia glauca Engl
Calophyllum lowii Hook. f.
Litsea nidularis Gamble
Shorea parvifolia Dyer.
Eugeissona utilis Becc.
Agathis borneensi Warb.
Litsea nidularis Gamble
Agathis borneensis Warb.
A Field Guide to the Permanent Sample Plots • 77
Table 6. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
Species
Dipterocarpus stellatus Vesque
Lithocarpus pulcher (King) Markgr.
Shorea beccariana Burck.
Dead
Syzygium sp. 5
Chisocheton sp.
Lithocarpus blumeanus (Korth.) Rehd.
Pentace laxiflora Merr.
Myristica villosa Warb.
Tarenna incerta Kds. et Val.
Palaquium calophyllum (T. & B.) Pierre
Dead
Shorea fallax Meijer
Madhuca magnifica H.J.Lam
Sterculia cordata Bl.
Shorea fallax Meijer
Mezzetia parviflora Becc.
Knema latericia Elmer
Dead
Artocarpus integer (Thunb.) Merr.
Mezzetia parviflora Becc.
Gymnacranthera contracta Warb.
Teijsmaniodendrum simplicifolium Merr.
Parishia maingayi Hook.f.
Shorea exelliptica Meijer
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
100
80
60
40
20
0
0
0
3
95
97
214
217
215
216
218
219
220
237
2 1
98
93 5
103
9
8
208
20
18
10 11
23
12
19
89
86
16
15 14
17
90
88
85
196
22
20
83
191
40
21
31
30
79
77 78
29
76
26
74
28
81
261
60
68
35
128
69
41
34
32 33
40
72
278
42
39
37
67
132
135
166
137
46
80
38
65
62
43
44
47
48
64 63
61
141
138
59
49
60
152
53
50
100
51 52
55
56
58
146
147
149
150
148
57
151
144
143
140
139
156
158
157
167
153
155 154
164
163
170
168
169
173
287
286
159
282
284
160
100
289 290
288
283
285
281
280
162 161
133 142
171
134
275
276
80
175
172
273
277
36
131
70 71
129
185 184
127
179
272
269
177
178 176
182
183
271
180
181
270
260
186
188
27
75
122
126
121
263
264
268
265 267
266
60
262
189
259
187
251
82
123
253
252
124 125
117
115
114
194
247
245
244
40
248
250
246
130
204
201
200
197
112
202
203
110
207 206
209
210
198
243
240
241 242
236 235
232
20
231
230
225
227 226
105
107
108
104
222
221
223
100
80
60
40
20
0
Figure 10. Distribution of Trees in Plot RIL07
Junglandaceae
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Apocynaceae
Annonaceae
Verbenaceae
Ulmaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Proteaceae
Polygalaceae
Oxalidaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
FAMILY
Anacardiaceae
Lauraceae
RIL07
78 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 79
Table 7. List of Trees in Plot RIL07
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Horsfiedla crassifolia (Hk.f.et Th.) Warb.
Oncosperma horridum Scheff.
Pholidocarpus sp
Dipterocarpus stellatus Vesque
Syzygium chlorantum (Duthie) Merr & Perry
Dipterocarpus lowii Hook.f.
Teijsmaniodendron simplicifolium Merr.
Lithocarpus pulcher (King) Markgr.
Lophopetalum subobovatum King
Dillenia grandiflora Wall ex Hook.f.
Shorea macroptera Dyer
Palaquium ferox H.J.Lam
Myristica beccarii Warb.
Vatica albiramis Sloot.
Koompassia malaccensis Maing. ex Benth.
Lithocarpus pulhcer (King) Markgr.
Shorea brunnescens P. S. Ashton
Ternstroemia aneura Miq.
Gironniera subaequalis Planch.
Zyzygium sp.
Palaquium calophyllum (T. & B.) Pierre
Anisoptera maegistocarpa Slooten
Tetramerista glabra Miq.
Dead
Pternandra galeata Cogn.
Callophyllum lowii Hook. f.
Shorea ovalis (Korth.) Bl.
Xanthophllum amoenum Chod.
Baccaurea edulis Merr.
Plectronia conferta (Korth.) Vall.
Swintonia glauca Engl.
Pouteia malaccensis (Clarke) Baehni.
Syzygium clhoranthum (Duthie) M.& P.
Pternandra galeata Cogn.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
80 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Horsfiedia crassifolia (Hk.f.et Th.) Warb.
Oncosperma horridum Scheff.
Syzygium grandis Wight.
Gymnacranthera contracta Warb.
Pternandra galeata Cogn.
Pholidocarpus sp.
Aporusa prainiana King ex Gage
Chionanthus ellipticus Bl.
Shorea ochracea Sym.
Quercus sp.
Shorea macroptea Dyer
Helicia petiolaris Benn.
Dipterocarpus stellatus Vesque
Tetramerista glabra Miq.
Dipterocarpus stellatus Vesque
Gluta wallichii (Hk.f.) Ding Hou
Dead
Artocarpus integer (Thunb.) Merr.
Shorea rubra Ashton
Canarium littorale Bl.
Sorea maxwelliana King
Dipterocarpus lowii Hk.f.
Shorea rubra Ashton
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Dillenia grandifolia Wall. ex Hk.f.
Dead
Dillenia grandifolia Wall. ex Hk.f.
Myristica beccarii Warb.
Shorea parvifolia Dyer
Gluta wallichii (Hkf.) Ding Hou
Gymnacranthera contracta Warb.
Pometia pinnata Forst.
Dead
Syzygium chloranthum (Duthie) M. & P.
A Field Guide to the Permanent Sample Plots • 81
Table 7. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Artocarpus anisophyllus Miq.
Syzygium sp.
Madhuca magnifica H.J.Lam
Cryptocarya tomentosa Bl.
Dacryodes incurvata (Engl.) H.J.Lam
Shorea exelliptica Meijer
Shorea macroptea Dyer
Pternandra galeata Cogn.
Syzygium grandis Wight.
Pimeleodendron papaveroides J.J.S.
Sageraea elliptica (DC.) Hk.f. ex Th.
Syzygium chloranthum (Duthie) M.& P.
Myristica beccarii Warb.
Koompassia malaccensis Maing ex Benth.
Dead
Vatica vinosa Ashton
Lophopetalum subobovatum King
Sandoricum emarginatum Hiern.
Teijsmaniodendron simplicifolium Merr.
Helicia petiolaris Benn.
Syzygium sp.5
Lophopetalum subobovatum King
Dyera costulata (Miq.) Hk.f.
Tetramerista glabra Miq.
Pternandra galeata Cogn.
Palaquium rostratum (Miq.) Burck
Shorea brunnescens P.S. Ashton
Shorea brunnescens P.S. Ashton
Palaquium ferox H.J.Lam
Gluta wallichii (Hk.f.) Ding Hou
Heritiera simplicifolia (Mart.) Kostrm.
Drypetes kikir Airy Shaw
Ternstroemia aneura Miq.
Baccaurea sumatrana M.A.
Shorea brunnescens P.S.Ashton
82 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Tetramerista glabra Miq.
Teijsmaniodendron simplicifolium Merr.
Syzygium sp.
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus stellatus Vesque
Palaquium sumatranum Burck
Shorea exelliptica Meijer
Castanopsis hypophoeniea (Seem.) Soepadmo
Dead
Tetramerista glabra Miq.
Madhuca malaccensis (Clarke) H.J.Lam
Shorea brunnescens P. S. Ashton
Teijsmaniodendron simplicifolium Merr.
Polyalthia glauca Boerl.
Pholidocarpus sp.
Dacryodes rostrata H.J.Lam
Gluta wallichii (Hk.f.) Ding Hou
Pholidocarpus sp.
Myristica beccarii Warb.
Shorea parvifolia Dyer
Shorea brunescens P.S Ashton
Gluta wallichii (Hk.f) Ding Hou
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
Teijsmaniodendron simplicifolium Merr
Endiandra coriacea Merr.
Dyera costulata (Miq.) Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Santiria griffithii (Hkf.) Engl.
Shorea macroptera Dyer
Shorea rubra P.S. Ashton
Dead
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea parvifolia Dyer
Shorea rubra P.S. Ashton
A Field Guide to the Permanent Sample Plots • 83
Table 7. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea rubra P.S. Ashton
Myristica villosa Warb.
Dyera costulata (Miq.) Hk.f.
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Shorea parvifolia Dyer
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus kunstleri (King) Warb.
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus lowii Hook.f.
Pholidocarpus sp.
Shorea maxwelliana King
Artocarpus integer (Thunb.) Merr
Gluta wallichii (Hk.f.) Ding Hou
Durio lanceolatus Mast.
Koompassia malaccensis Maing. ex Benth.
Dead
Engelhardia serrata Bl.
Shorea exelliptica King
Dacryodes incurvata H.J.Lam
Shorea maxwelliana King
Mezzetia parviflora Becc.
Gluta wallichii (Hk.f.) Ding Hou
Dialium maingayi Baker
Shorea macroptera Dyer
Mezzetia parviflora Becc.
Syzygium sp.
Mangifera magnifica Kochumen
Sindora leiocarpa Backer ex de Wit
Gluta wallichii (Hk.f.) Ding Hou
Memecylon myrsinoides Blume
Cryptocarya tomentosa Bl.
Horsfieldia crassifolia (Hk.f. et Th.) Warb.
Sindora leiocarpa Backer ex de Wit
Shorea parvifolia Dyer
Shorea pauciflora King
Pometia pinnata Forst.
84 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Buchanania arborescens Bl.
Myristica villosa Warb.
Diospyros oblonga Wall. ex G. Don
Shorea maxwelliana King
Vatica sp.
Sandoricum emarginatum Hiern.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Lophopetalum subobovatum King
Litsea grandis (Wall. ex Nees) Hook.f.
Dead
Pternandra galeata Cogn.
Calophyllum lowii Hk.f.
Mangifera blommestenii Kosterm.
Buchanania arborescens Bl.
Shorea parvifolia Dyer
Pternandra galeata Cogn.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Teijsmaniodendron simplicifolium Merr.
Dead
Dipterocarpus stellatus Vesque
Helicia fuscotomentosa Suesseng.
Pternandra galeata Cogn.
Dacryodes rugosa (Bl.) H. J. Lam
Sarcotheca diversifolia (Miq.) Hall.f.
Macaranga conifera (Zoll.) M.A.
Shorea parvifolia Dyer
Pometia pinnata Forst.
Koompassia malaccensis Maing. ex Benth.
Teijsmaniodendron simplicifolium Merr.
Vatica sp.
Pometia pinnata Forst.
Knema laterifolia Warb.
Shorea maxwelliana King
Tetramerista glabra Miq.
Dacryodes incurvata H.J.Lam
Shorea maxwelliana King
A Field Guide to the Permanent Sample Plots • 85
Table 7. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Baccaurea sumatrana M.A.
Dipterocarpus lowii HK.f.
Heritiera javanica (Bl.) Kosterm.
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Dead
Vatica sp.
Myristica iners Bl.
Litsea grandis (Wall.ex Ness) Hook.f
Mezzetia parviflora Becc.
Pometia pinnata Forst.
Mangifera blommestenii Kosterm.
Shorea maxwelliana King
Artocarpus integer (Thunb.) Merr.
Memecylon myrsinoides Blume
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea maxwelliana King
Bacaurea sumatrana M.A.
Shorea parvifolia Dyer
Sandoricum emarginatum Hiern.
Syzygium sp.
Calophyllum lowii Hk.f.
Tetramerista glabra Miq.
Shorea pauciflora King
Shorea exelliptica Meijer
Dead
Dillenia grandiflora Wall.ex Hk.f
Sarcotheca diversifolia (Miq.) Hall.f
Lophopetalum subobovatum King
Shorea venulosa Wood.ex Meijer
Gironniera subaequalis Planch.
Artocarpus integer (Thynb.) Merr.
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus lowii Hk.f.
Syzygium sp.
86 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
Species
Dillenia grandifolia Wall.ex Hk.f.
Shorea pauciflora King
Gymnacranthera contracta Warb.
Scaphium macropodum (Miq.) Beumée ex Heyne
Palaquium cochlearifolium van Royen
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Teijsmaniodendron simplicifolium Merr.
Gluta wallichii (Hk. f.) Ding Hou
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Dillenia grandifolia Wall.ex Hk.f
Gymnacranthera contracta Warb.
Dead
Shorea maxwelliana King
Shorea exelliptica Meijer
Shorea rubra Ashton
Shorea maxwelliana King
Dead
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Pternandra galeata Cogn.
Tentramerista glabra Miq.
Diospyros dayakensis Bakh.
Pholidocarpus sp.
Buchanania arborescens (Bl.) Bl.
Ctenolophon parvifolius Oliv.
Dipterocarpus lowii Hook.f
Shorea rubra P.S.Ashton
100
80
60
40
20
0
0
0
1
103
109
108
111
3
5
106 105
104
110
119 123
9
6
7 8
4
96
13
99
102
107
20
10
11
12
93
129
217
220
259
17
16
22
19
21
85
131
132
134
133
92
14 15
91
90
89
88
128 86
87
127
126
94
97
95
98
125
218
124
101
100
232
231
230
40
80
79
40
20 28
81
77
29
24
75
25
23
74
73
30
32
72
150
149
148
147
141
138
140
146
144
145
209
272
271
66
31
68 69
67
71
151
152
60
70
65
155
154
60
204
202
273
274
203
270
205
208
207 206
136
137
139
27
142
135
216
215
214
212
225
263
262
210
269
261
268
260
223
221
224
222
258
257
253
266
265 267
254
256
255
229 228
249
250
121
120
122
118
112 113
114
115
233
234
235
236
239 238
243
244 242
241
247
246
20
251 252
33
61
163
153
162
158
159
167
40
60
34
62
160
161
156
199
200
38
39
63
164
166
35
37
36
42
80
58
172
173
59
41
197
193
57
56
174
175
100
43
44
47
100
49
50
52
54
55
180
48
53
178
179
181
183
182 184
186
189 185
187
45 46
177
176
190
191
188
293
194 192
195
292
196
171
286
291
168 169
198
201
281 285
277 282
284
280
279
275
287
289 288
276
80
100
80
60
40
20
0
Figure 11. Distribution of Trees in Plot RIL08
Lauraceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Aquifoliaceae
Apocynaceae
Annonaceae
Verbenaceae
Ulmaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rosaceae
Rhizophoraceae
Polygalaceae
Podocarpaceae
Oxalidaceae
Oleaceae
Ochnaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
FAMILY
Anacardiaceae
Lecythidaceae
RIL08
A Field Guide to the Permanent Sample Plots • 87
88 • A Field Guide to the Permanent Sample Plots
Table 8. List of Trees in Plot RIL08
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Syzygium chloranthum (Duthie) Merr. & Perry
Dead
Lophopetalum subobovatum King
Elaeocarpus petiolata Wall.
Shorea venulosa Wood ex Meijer
Ternstroemia aneura Miq.
Tetramerista glabra Miq.
Shorea maxwelliana King
Pouteria malaccenis (Clarke) Baehni.
Shorea venulosa Wood ex Meijer
Shorea maxweliana King
Pternandra galeata Cogn.
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Calophyllum lowii Hk.f.
Shorea rubra Ashton
Dead
Memecylon myrsinoides Blume
Baccaurea sumatrana M.A
Santiria laevigata Bl.
Gluta wallichii (Hk.f.) Ding Hou
Koompassia malaccensis Maing. ex Benth.
Myristica beccarii Warb.
Chionanthus pubicalyx (Ridl.) Kiew.
Unidentified
Dialium maingayi Baker
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Lophopetalum subobovatum King
Shorea venulosa Wood ex Meiyer
Gymnacranthera contracta Warb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Lophopetalum subobovatum King
Pouteria malaccenis (Clarke) Baehni.
Shorea venulosa Wood ex Meijer
Vatica umbonata (Hook.f.) Burck
Shorea rubra Ashton
A Field Guide to the Permanent Sample Plots • 89
Table 8. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Calophyllum lowii Hk.f.
Madhuca palembanica (Miq.) P.C.Yii & P.Chai
Calophyllum pulcherrimum Wall. ex Choisy
Mezzetia parviflora Becc.
Shorea rubra Ashton
Shorea rubra Ashton
Syzygium sp. B
Shorea rubra Ashton
Lophopetalum subobovatum King
Litsea firma Bl.
Prunus arborea (Bl.) Kalkm.
Endiandra coriacea Merr.
Shorea venulosa Wood ex Meijer
Dead
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Elaeocarpus longipetiolatus Merr.
Elaeocarpus longipetiolatus Merr.
Elaeocarpus petiolata Wall.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Swintonia glauca Engl.
Gironniera subaequalis Planch.
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum lowii Hk.f.
Dillenia grandifolia Wall. ex Hk.f.
Syzygium bankensis (Hassk.) Merr. & Perry
Buchanania arborescens (Bl.) Bl.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Aglaia ganggoo Miq.
Shorea venulosa Wood ex Meijer
Palaquium rostratum (Miq.) Burk.
Madhuca erythrophylla (K.ex G.) H.J.Lam
Actinodaphne borneensis Meissn.
Baccaurea sumatrana M.A
Shorea venulosa Wood ex Meijer
Calophyllum lowii Hk.f.
Anisophyllea corneri Ding Hou
90 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Gluta wallichii (Hk.f.) Ding Hou
Pternandra galeata Cogn.
Petrenandra galeata Cogn.
Dead
Syzygium incarnatum (Elmer) Merr. & Perry
Lycania splendens (Korth.) Prance
Madhuca erythrophylla (K. & G.) H.J.Lam
Unidentified
Unidentified
Unidentified
Teijsmaniodendron simplicifolium Merr.
Calophyllum lowii Hk.f.
Garcinia rostrata T. et B.
Lophopetalum subobovatum King
Madhuca malaccensis (Clarke) H.J.Lam
Shorea maxwelliana King
Shorea rubra Ashton
Gluta wallichii (Hk.f.) Ding Hou
Lophopetalum subobovatum King
Calophyllum lowii Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Scaphium macropodum (Miq.) Buemée ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Mezzetia parviflora Becc.
Gluta wallichii (Hk.f) Ding Hou
Gironniera subaequalis Planch.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata Bl.
Myristica beccarii Warb.
Shorea rubra Ashton
Mangifera blommesteinii Kostrm.
Dipterocarpus lowii Hk.f.
Swintonia glauca Engl.
Santiria laevigata Bl.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea exelliptica Meijer
A Field Guide to the Permanent Sample Plots • 91
Table 8. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Dialum maingayi Baker
Ctenolophon parvifolius Oliv.
Shorea exelliptica Meijer
Koompassia malaccensis Maing ex Benth.
Gymnacranthera contracta Warb.
Shorea exelliptica Meijer
Shorea venulosa Wood ex Meijer
Cleistanthus bakonensis Airy Shaw
Barringtonia scortechinii King
Dacryodes rostrata H.J.Lam
Querqus gemmeliflora Bl.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Palaquium calophyllum ( T. et B. ) Pierre
Pholidacarpus sp.
Sageraea alliptica (DC.) Hk.f. et Th.
Sandoricum emarginatum Hiern
Dacryodes incurvata (Engl.) H.J.Lam
Dead
Calophyllum lowii Hk.f.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Artocarpus anisophyllus Miq.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Myristica beccarii Warb.
Calophyllum lowii Hk.f.
Madhuca erithrophylla ( K & G.) H.J.Lam
Calophyllum lowii Hk.f.
Licania spendens (Korth.) Prance
Syzygium ochneocarpa Merr.
Shorea atrihervosa Sym
Dipterocarpus lowii Hk.f.
Dead
Calophyllum pulcherrimumWall. ex Choisy
Shorea rubra Ashton
Elaeocarpus longipetiolatus Merr.
Madhuca erythrophylla ( K. ex G. ) H.J.Lam
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
92 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea atrinervosa Sym.
Syzygium incarnatum (Elmer) Merr. & Perry
Memecylon sp.
Shorea venulosa Wood ex Meijer
Shorea exelliptica Meijer
Sarcotheca diversifolia (Miq.) Hall.f.
Agathis borneensis Warb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Dead
Garcinia penangiana Pierre
Pholidocarpus sp.
Shorea brunnesens Ashton
Shorea brunnesens Ashton
Lophopetalum subobovatum King
Shorea venulosa Wood ex Meijer
Gluta wallichii (Hk.f.) Ding Hou
Dead
Dipterocarpus lowii Hk.f
Tetramerista glabra Miq.
Alstonia pneumatophora Baker
Baccaurea bracteata M.A.
Dead
Tetramerista glabra Miq.
Tetramerista glabra Miq.
Ctenolophon parvifolius Oliv.
Gluta wallichii (Hk.f.) Ding Hou
Gironniera subaequalis Planch.
Dipterocarpus stellatus Vesque
Myristica beccarii Warb.
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes incurvata (Engl.) H.J.Lam
Gluta wallichii (Hk.f.) Ding Hou
Myristicca beccarii Warb.
Mezzetia parviflora Becc.
Gluta walichii (Hk.f.) Ding Hou
Dehaasia firma Bl.
Teijsmaniodendron simpliciafolium Merr.
A Field Guide to the Permanent Sample Plots • 93
Table 8. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Gomphia serrata Gaertn.
Tetramerista glabra Miq.
Dipterocarpus lowii Hook.f.
Tetramerista glabra Miq.
Dipterocarpus stellatus Vesque
Calophyllum lowii Hk.f.
Syzygium grandis Wight.
Pholidocarpus sp.
Madhuca mallacensis (Clarke) H.J.Lam
Lithocarpus cantleyanus (King ex Hk.f.) Rehd .
Buchanania arborescers (Bl.) Bl.
Garcinia maingayi Hook.f.
Gluta wallichii (Hk.f ) Ding Hou
Chionantus ellipticus Bl.
Tetramerista glabra Miq.
Diospyros sp.
Tetramerista glabra Miq.
Dehaasia firma Bl.
Garcinia nervosa Miq.
Diospyros oblonga Wall. ex G. Don
Ilex sp.
Tetramerista glabra Miq.
Dead
Durio testudinarum Becc.
Dead
Tetramerista glabra Miq.
Koompassia malaccensis Maing ex Benth.
Baccaurea sumatrana M. A.
Shorea venulosa Wood ex Meijer
Oncosperma horridum Scheff.
Dead
Madhuca palembanica (Miq.) P.C.Yii & P.Chai
Lithocarpus benettii (Miq.) Rehd.
Ctenolophon parvifolius Oliv.
94 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Callophylum lowii Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Prunus arboreus (Bl.) Kalkm.
Dead
Dead
Lithocarpus cantleyanus(King & Hk.f.) Rehd.
Ctenolophon parvifolius Oliv.
Shorea venulosa Wood ex Meijer
Shorea exelliptica Meijer
Syzygium chloranthum (Duthie) Merr & Perry
Dipterocarpus lowii Hk.f.
Calophyllum pulcherrimum Wall. ex Choisy
Shorea venulosa Wood ex Meijer
Dypetes kikir Airy Shaw
Dead
Baccaurea sumatrana M.A.
Canarium littorale Bl.
Dead
Paratocarpus forbesii King
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea venulosa Wood ex Meijer
Baccaurea sumatrana M.A
Dead
Syzygium ochneocarpa Merr.
Agathis borneensis Warb.
Artocarpus integer (Thunb.) Merr.
Baccaurea pendula Merr.
Litsea nidularis Gamble
Artocarpus anisophyllus Miq.
Podocarpus neriiforlius D.Don.
Koompassia malancensis Maing. ex Benth.
Xanthophyllum rufum Bth.
Artocarpus anisophyllus Miq.
Tetramerista glabra Miq.
Pometia pinnata Forst.
Syzygium acutangulum K. Schum.
Shorea brunnescens P.S.Ashton
A Field Guide to the Permanent Sample Plots • 95
Table 8. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
Species
Syzygium chlorontum (Duthie) M. & P.
Shorea venulosa Wood ex Meijer
Syzygium chloranytum (Duthie) M. & P.
Archidendron fagifolum (Bl. ex Miq.) Nielsen
Unidentified
Mezzetia parvifolra Becc.
Palaquium sumatranum Burck
Litsea nidularis Gamble
Dialium maingayi Baker
Shorea venulosa Wood ex Meijer
Depterocarpus lowii Hook.f.
Dipterocarpus stellatus Vesque
Litsea nidularis Gamble
Litsea nidularis Gamble
Durio lanceolatus Mast.
Syzygium sp.
Shorea ochracea Sym.
Podocarpus neriifolius D.Don
Prunus arborea (Bl.) Kalkm.
Ctenolophon parvifolius Oliv.
Tetramerista glabra Miq.
Lophopetalum subobovatum King
Gluta wallichii ( Hk.f.) Ding Hou
Dead
Shorea exelliptica Meijer
Tetramerista glabra Miq.
Tetramerista glabra Miq.
Trigonapleura malayana
Tetramerista glabra Miq.
Xylopia altissima Boerl.
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Dipterocarpus lowii Hook.f.
Mesua congestiflora P.F.Stevens
Dipterocarpus lowii Hook.f.
96 • A Field Guide to the Permanent Sample Plots
Table 9. List of Trees in Plot RIL09
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Palaquium cochlearifolium van Rpoyen
Palaquium cochlearifolium van Royen
Shorea venulosa Wood ex Meijer
Agathis borneenis Warb.
Calophyllum biflorum H. & W. Smith
Shorea parvifolia Dyer
Dead
Palaquium cochlearifolium van Royen
Endiandra coriacea Merr.
Dead
Myristica beccarii Warb.
Shorea maxwelliana King
Dead
Shorea maxwelliana King
Memecylon myrsinoides Bl.
Baccaurea javanica M.A.
Shorea maxwelliana King
Cleistanthus bakonensis Airy Shaw
Baccaurea javanica M.A.
Hydnocarpus kunsleri (King )Warb.
Pouteria malaccensis (Clarke) Baehni.
Dead
Pouteria malaccensis (Clarke) Baehni.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Myristica beccarii Warb.
Mangifera swintonioides Kosterm.
Buchanania arborescens Bl.
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea maxwelliana King
Anisoptera megistocarpa Slooten
Shorea exelliptica Meijer
A Field Guide to the Permanent Sample Plots • 97
Table 9. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dialium maingayi Baker
Mezzetia parviflora Becc.
Myristica beccarii Warb
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata Bl.
Ternstroemia aneura Miq.
Shorea maxwelliana King
Shorea parvifolia Dye
Sindora leiocarpa Backer ex De Wit
Calophyllum pulcherrimum Wall ex Choisy
Pouteria malaccensis (Clarke) Baehni.
Shorea exelliptica Meijer
Syzygium albidiramea Merr.
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum lowii Hk.f.
Dead
Dead
Dehaasia incrassata (Jack.) Kosterm.
Madhuca erythrophylla (K.ex G.) H.J.Lam
Shorea rubra Ashton
Hydnocarpus kunstleri (King) Warb.
Shorea maxwelliana King
Shorea machroptera Dyer
Shorea beccarii Burck
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum lowii Hk.f.
Syzygium chloranthum (Duthie) Merr & Perry
Syzygium clavimyrtus K. et V.
Gymnacranthera contracta Warb.
Agathis borneensis Warb.
Shorea atrinervosa Sym
Myristica iners Bl.
Calophyllum pulcherrimum Wall. ex Choisy
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck
Shorea parvifolia Dyer
Shorea atrinervosa Sym
98 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Sindora leiocarpa Baker
Shorea maxwelliana King
Calophyllum pulcherrimum Wall. ex Choisy.
Scaphium macropodum (Miq.) Beumee
Litsea firma Bl.
Anisoptera megistocarpa Slooten
Dead
Gymnacranthera contracta Warb.
Shorea rubra Ashton
Shorea rubra Ashton
Dead
Diptetocarpus lowii Hk.f.
Gluta wallichii (Hk.f. ) Ding Hou
Artocarpus anisophylus Miq
Syzygium caudatilimbum Merr. & Perry
Hydnocarpus kunstleri (King & Hkl.f.) Rehd.
Pouteria malaccensis (Clarke) Baehni.
Mezzetia parviflora Becc.
Dialium sp.
Myristica beccarii Warb.
Gluta wallichii (Hk.f.) Ding Hou
Xanthophyllum sp.
Shorea exelliptica Meijer
Cleistanthus ferrugineus M.A.
Shorea maxwelliana King
Shorea rubra Ashton
Shorea rubra Ashton
Dead
Durio lanceolatus Mast.
Castanopsis javanica (Bl.) A.DC.
Shorea parvifolia Dyer ssp parvifolia
Shorea beccarriana Burck
Memecylon myrsinoides Blume
Endiandra coriacea Merr.
Gonystilus forbesii Gilg.
Shorea beccariana Burck
Swintonia glauca Engl.
A Field Guide to the Permanent Sample Plots • 99
Table 9. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Durio lanceolatus Mast.
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King & Hk.f.)Rehd.
Myristica beccarii Warb.
Dead
Shorea parvifolia Dyer ssp parvifolia
Pouteria malaccensis (Larke) Baehni.
Syzygium ochneocarpa Merr.
Myristica beccarii Warb.
Pouteria malaccensis (Larke) Baehni.
Calophyllum lowii Hk.f.
Shorea beccariana Burck
Dead
Shorea maxwelliana king
Dipterocarpus lowii Hk.f.
Santiria laevigata Bl.
Shorea agami Ashton
Agathis borneesis Warb.
Hydnocarpus kunstleri (King) Warb.
Agathis borneensis Warb.
Palaquium cochlearifolium van Royen
Koordersiodendron pinnatum (Blanco) Merr.
Shorea venulosa Wood ex Meijer
Vatica umbonata (Hook.f.) Bl.
Sarcotheca diversifolia (Miq.) Hall.f.
Syzygium cf. angustifolius (Miq.) Hats.
Shorea macroptera Dyer
Amoora lanceolatus Heim.
Durio lanceolatus Mast.
Parashorea smithiesii Wyatt-Smith ex Asht.
Elaeocarpus polystachys Wall.
Licania splendens (Korth.) Prance
Shorea agami Ashton
Dead
Dacryodes rostrata H.J.Lam
Diospyros oblonga Wall ex G. Don
Madhuca palembanica (Miq.) P.Y.Yii & P.C.
100 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Syzygium grandis Wight.
Sindora velutina Backer ex De wit
Dead
Gymnacranthera contracta Warb.
Shorea agami Ashton
Shorea parvifolia Dyer ssp parvifolia
Swintonia glauca Engl.
Dead
Lithocarpus cantleyanus (King & Hk.f.)Rehd.
Lithocarpus cantleyanus (King & Hk.f.)Rehd.
Myristica beccarii Warb.
Lithocarpus cantleyanus (King & Hk.f.)Rehd.
Lithocarpus cantleyanus (King & Hk.f.)Rehd.
Dead
Dead
Gomphia serrata (Gaertn.) Kanis
Vatica umbonata (Hook.f.) Bl.
Agathis borneensis Warb.
Shorea venulosa Wood ex Meijer
Syzygium bankensis (Hassk.) Merr & Perry
Agathis borneensis Warb.
Gluta wallichii (Hk.f.)Ding Hou
Gymnacranthera contracta Warb.
Dipterocarpus lowii Hk.f.
Vatica umbonata (Hook.f.) Bl.
Agathis borneensis Warb.
Santiria laevigata Bl.
Agathis borneensis Warb.
Aglaia ganggoo Miq.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Myristica beccarii Warb.
Cleistanthus ferrugineus M.A.
Shorea maxwelliana King
Syzygium angustifolius (Miq.) Hats.
Gimnacranthera contracta Warb
Shorea rubra Ashton
A Field Guide to the Permanent Sample Plots • 101
Table 9. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Myristica beccarii Warb.
Dead
Cleistanthus ferrugineus M.A.
Shorea rubra Ashton
Gluta wallichii (Hk.f.) Ding Hou
Memecylon myrsinoides Blume
Calophyllum lowii Hk.f.
Palaquium cochlearifolium van Royen
Shorea venulosa Wood ex Meijer
Koompassia malaccensis Maing ex Benth.
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum pulcherrimum Wall. ex Choisy.
Swintonia glauca Engl.
Santiria griffithii (Hk.f.) Engl.
Shorea maxwelliana King
Baccaurea sumatrana M. A.
Syzygium sp.
Gironniera subaequalis Planch.
Syzygium sp.
Dacryodes rugosa H.J.Lam
Gomphia serrata (Gaertn.) Kanis
Castanopsis motleyanus King
Shorea venulosa Wood ex Meijer
Gluta wallichii (Hk.f.) Ding Hou
Elaeocarpus cf. petiolatus Merr.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Gironniera subaequalis Planch.
Diospyros sp.
Endiandra sp.
Baccaurea sumatrana M. A.
Baccaurea sp.
Dead
Licania splendens (Korth.) Prance
Shorea maxweliana King
Myristica becarii Warb.
Heritiera javanica (Bl.) Kosterm.
102 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Madhuca malaccensis (Clarke) H.J.Lam
Gomphia serrata (Gaertn.) Kanis
Crypteronia griffithii Clarke
Cratoxylon arborescens
Timonius borneensis Val.
Dipterocarpus lowii Hk.f.
Shorea parvifolia Dyer
Shorea exelliptica Meijer
Adinandra borneensis Kob.
Madhuca malaccensis (Clarke) H.J.Lam
Shorea maxwelliana King
Madhuca palembanica (Miq.) P.Y.Yii & P.C.
Artocarpus integer (Thunb.) Merr.
Calophyllum lowii Hk.f.
Shorea beccariana Burck
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Licania splendens (Korth.) Prance
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea macroptera Dyer
Adinandra borneesnsis Kob.
Myristica beccarii Warb.
Vatica albiramis Sloot.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea maxwelliana King
Dysoxylum hexandrum Merr.
Xanthophyllum amoenum Chod.
Shorea maxwelliana King
Cleistanthus cf. ferrugineus M.A.
Agathis borneensis Warb.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Madhuca palembanica (Miq.) P.Y.Yii & P.C.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Licania splendens (Korth.) Prance
Xanthophyllum amoenum Chod.
A Field Guide to the Permanent Sample Plots • 103
Table 9. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Cleistanthus ferrugineus M.A.
Hydnocarpus kunstleri (King) Warb.
Gymnacranthera contracta Warb.
Syzygium grandis Wight.
Shorea parvifolia Dyer
Shorea maxwelliana King
Syzygium chloranthum (Duthie) Merr & Perry
Palaquium cochlearifolium van Royen
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Artocarpus anisophyllus Miq.
Dead
Baccaurea sumatrana M.A.
Licania splendens (Korth.) Prance
Cleistanthus ferrugineus M.A.
Ternstroemia aneura Miq.
Santiria laevigata Bl.
Atuna excelsa (Benth.) Kosterm.
Lithocarpus pulcher (King) Markgr.
Santiria laevigata Bl.
Calophyllum lowii Hk.f.
Parashorea smithiesii Wyatt-Smith ex Asht.
Shorea parvifolia Dyer
Campnosperma auriculata Hk.f.
Baccaurea sumatrana M. A.
Koompassia malaccensis Maing. ex Benth.
Lithocarpus ewyckii (Korth.) Rehd.
Baccaurea kunstleri King
Koompassia malaccensis Maing. ex Benth.
Tetramerista glabra Miq.
Syzygium sp.
Gironniera nervosa Planch.
Calophyllum lowii Hk.f.
Shorea pauciflora King
Hydnocarpus kunstleri (King ) Warb.
Adinandra borneensis Kob.
Shorea pauciflora King
104 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
307
308
309
310
311
312
313
314
315
316
317
318
319
320
321
322
323
324
325
326
327
328
329
330
331
332
333
Species
Shorea beccarriana Burck
Macaranga hosei King ex Hook.f.
Oncosperma horridum Scheff.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Shorea parvifolia Dyer
Shorea macroptera Dyer
Syzygium grandis Wight.
Baccaurea kunstleri King
Syzygium grandis Wight.
Cratoxylon arborescens Bl.
Santiria laevigata Bl.
Shorea parvifolia Dyer
Knema latifolia Warb.
Hydnocarpus kunstleri (King ) Warb.
Adinandra borneensis Kob.
Baccaurea sumatrana M.A.
Shorea pauciflora King
Anisoptera megistocarpa Slooten
Shorea beccarriana Burck
Syzygium grandis Wight.
Oncosperma horridum Scheff
Lithocarpus ewyckii (Korth.) Rehd.
Shorea fallax Meijer
Oncosperma horridum Scheff.
Dehaasia incrassata Bl
Gironniera subaequalis Planch
Macaranga hosei King ex Hook.f.
Lithocarpus pulcher (King) Markgr.
Gironniera subaequalis Planch.
Drospyros
Madhuca erythrophylla (K.ex G.) H.J.Lam
Glochidion borneense Boerl.
Macaranga conifera (Zoll.) M.A.
Baccaurea sumatrana M.A.
Dialium sp.
Adinandra borneensis (bulu)
Timonius borneensis Val.
100
80
60
40
20
0
0
0
256
81
230
18
49
25
192
38
4
79
87
269
37
20
51
166
235
155
116
90
204
40
187
167
247
21
56
188
71
94
135
162
40
258
246
171
14
196
186
257
54
33 82
234
244
245
201
36
73
75
209
199
268
1
168
278
273
53
203
5
279
92
91
117
134
16
132
131
169
267
118
226
170
161
185
20
237
78
13
277
86
119
227
128
229 130
249
189
123
219
85
121
241
129
2
265
19
59
57
58
236
250
214
22
262
44
35
238
159
60
34
3
266
239
115
61
215
181
149
67
10
60
259
24
9
113
111
289
143
281
158
139 254
287
17 93
29
243
60
11
197
261
248
148
222
112
32
66
182
80
272
202
45
282
151
208
47
150
99
190
12
107
147
191
280
23
108
145
144
95
20 96
140
164
80
100
175
64
260
63
216
30
31
103
251
48
146
255
100
100
80
60
40
20
0
Figure 12. Distribution of Trees in Plot RIL10
Fagaceae
Fabaceae
Euphorbiacea
Ebenaceae
Dipterocarpaceae
Dilleniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Araucariaceae
Apocynaceae
Annonaceae
Ulmaceae
Thymelaeaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rosaceae
Polygalaceae
Oleaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Linaceae
Lauraceae
Junglandaceae
FAMILY
Anacardiaceae
Flacourtiaceae
RIL10
A Field Guide to the Permanent Sample Plots • 105
106 • A Field Guide to the Permanent Sample Plots
Table 10. List of Trees in PlotRIL10
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Palaquium cochlearifolium van Royen
Myristica maxima Warb.
Mezzetia parvifolia Becc.
Shorea exelliptica Meijer
Dialium laurinum Backer
Shorea parvifolia Dyer
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Syzygium sp.
Calophyllum lowii Hook.f.
Beilscmiedia madang Bl.
Syzygium chloranthum (Duthie) Merr. & Perry
Shorea exelliptica Meijer
Prunus arborea (Bl.) Kalkman
Dead
Swintonia glauca Engl.
Shorea beccariana Burck.
Parastemon urophyllus (A.DC.) A.DC.
Agathis borneensis Warb.
Ctenolophon parvifolius Oliv.
Lithocarpus cantleyanus (Kinq. ex HK.f.) Rehd.
Madhuca erythrophylla (K. & G.) H.J.Lam
Aphanamixis grandifolia Bl.
Shorea maxwelliana King
Xanthophyllum flavescens Roxb.
Vatica albiramis Slooten
Koompassia malaccensis Maing. ex Benth.
Vatica albiramis Slooten
Shorea beccariana Burck.
Licania splendens (Korth.) Prance
Hydnocarpus gracilis (v.Sl.) Sleum.
Mangifera swintonioides Kosterm.
Shorea macroptera Dyer
Palaquium cochlearifolium van Royen
Dyera costulata (Miq.) Hk.f.
Shorea beccariana Burck.
Shorea exelliptica Meijer
A Field Guide to the Permanent Sample Plots • 107
Table 10. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Calophyllum lowii Hook.f.
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Artocarpus integer (Thunb.) Merr.
Unidentified
Agathis borneensis Warb.
Pometia pinnata Fost.
Myristica maxima Warb.
Shorea parvifolia Dyer
Baccaurea puberula Merr.
Engelhardia serrata Bl.
Shorea parvifolia Dyer
Castanopsis motleyana King
Dead
Dipterocarpus stellatus Vesque
Gymnacrantera contracta Warb.
Palaquium cochlearifolium van Royen
Aphanamixis cf. grandifolia Bl.
Unidentified
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Shorea parvifolia Dyer
Myristica villosa Warb.
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Baccaurea sumatrana M.A.
Dead
Dipterocarpus stellatus Vesque
Pternandra galeata (Cogn.) Ridley
Agathis borneensis Warb.
Scaphium macropodum (Miq.) Beumee
Artocarpus integer (Thunb.) Merr.
Pimeleodendron papaveroides J.J.Sm.
Baccaurea sumatrana M.A.
Dead
Shorea brunnescens Ashton
Licania splendens (Korth.) Prance
Madhuca sp.
Agathis borneensis Warb.
108 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Shorea venulosa Wood ex Meijer
Shorea brunnescens Ashton
Mangifera sp.
Shorea macroptera Dyer
Syzygium attenuatum (Miq.) Merr. & Perry
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Myristica beccarii Warb.
Dipterocarpus stellatus Vesque
Shorea beccarriana Burck.
Baccaurea sumatrana M.A.
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum lowii Hook.f.
Hydnocarpus sp.
Shorea beccariana Burck.
Calophyllum lowii Hook.f.
Shorea agamii Ashton
Shorea pinanga Scheff.
Memecylon myrsinoides Blume
Baccaurea sumatrana M.A.
Palaquium cochlearifolium van Royen
Shorea beccariana Burck.
Dipterocarpus stellatus Vesque
Calophyllum pulcherrimum Wall. ex Choisy
Calophyllum lowii Hook.f.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Dead
Dipterocarpus stellatus Vesque
Syzygium caudatilimbum (Merr.) Merr & Perry
Lithocarpus sp.
Dead
Dead
Palaquium cochlearifolium van Royen
Shorea maxweliana King
Myristica beccarii Warb.
Calophyllum lowii Hook.f.
Myristica beccarii Warb.
A Field Guide to the Permanent Sample Plots • 109
Table 10. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea rubra Ashton
Calophyllum lowii Hook.f.
Agathis borneensis Warb.
Shorea venulosa Wood ex Meijer
Licania splendens (Korth.) Prance
Hydnocarpus kunstleri (King ) Warb.
Calophyllum lowii Hk.f.
Durio lanceolatus Mast.
Dead
Shorea maxwelliana King
Gonystylus forbesii Gilg.
Shorea maxwelliana King
Castanopsis javanica (Bl.) A.DC.
Shorea beccariana Burck
Dead
Myristica beccarii Warb.
Gluta wallichii (Hk.f) Ding Hou
Gonystylus forbesii Gilg.
Calophyllum lowii Hk.f.
Dialium maingayi Baker
Scaphium macropodum (Miq) Beumée ex Heyne
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Artocarpus anisophyllus Miq.
Gluta wallichii (Hk.f) Ding Hou
Shorea atrinervosa Sym.
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Gluta wallichii (Hk.f ) Ding Hou
Shorea venulosa Wood ex Meijer
Beilschmiedia madang Bl.
Dead
Shorea parvifolia Dyer
Gonystylus forbesii Gilg.
Calophyllum lowii Hk.f
Shorea atrinervosa Sym
Gluta wallichii (Hk.f) Ding Hou
Syzygium bankensis (Hassk) Merr & Perry
Calophyllum lowii Hk.f
110 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea maxwelliana King
Licania splendens (Korth.) Prance
Palaquium cochlearifolium van Royen
Shorea rubra Ashton
Dead
Dead
Canarium littorale Bl.
Tenstroemia aneura Miq.
Dead
Artocarpus anisophyllus Miq.
Palaquium cochlearifolium van Royen
Castanopsis motleyana King
Gluta wallichii (Hk.f) Ding Hou
Diospyros oblonga Wall. ex G. Don
Dipterocarpus stellatus Vesque
Gluta wallichii (Hk.f) Ding Hou
Gironniera subaequalis Planch.
Gymnacranthera contracta Warb.
Swintonia glauca Engl.
Dyera costulata (Miq) Hk.f.
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Dipterocarpus lowii Hook.f
Dipterocarpus stellatus Vesque
Endiandra coriacea Merr.
Aporusa anntenifera Airy Shaw
Shorea exelliptica Meijer
Shorea macroptera Dyer
Dipterocarpus stellatus Vesque
Llithocarpus cantleyanus (King & Hk.f) Rehd.
Hydnocarpus kunstleri (King) Warb.
Santiria laevigata Bl.
Shorea venulosa Wood ex Meijer
Santiria griffithii (Hk.f) Engl.
Shorea beccariana Burck
Gymnacranthera contracta Warb.
Gironniera subaequalis Planch.
A Field Guide to the Permanent Sample Plots • 111
Table 10. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Calophyllum lowii Hk.f.
Dillenia grandifolia Wall ex Hk.f.
Prunus arborea (Bl.) Kalkm.
Shorea maxwelliana King
Gluta wallichii (Hk.f) Ding Hou
Hydnocarpus kunstleri (King) Warb.
Shorea maxwelliana King
Dead
Dead
Agathis borneensis Warb.
Dipterocarpus stellatus Vesque
Lophopetalum subobouatum King
Dipterocarpus stellatus Vesque
Shorea exelliptica Meijer
Dipterocarpus lowii Hook.f
Shorea venulosa Wood ex Meijer
Cleistanthus bakonensis Airy Shaw
Baccaurea sumatrana M.A
Dipterocarpus lowii Hook.f.
Dead
Dead
Dead
Shorea maxwelliana King
Syzygium bankensis (Hassk.) Merr. & Perry
Myristica beccarii Warb.
Pimeleodendron papaveroides J.J.S.
Atuna excelsa (Jack) Kosterm.
Myristica beccarii Warb.
Sindora leiocarpa Backer ex de Wit.
Palaquium cochlearifolium van Royen
Scaphium macropodum (Miq) Beumee ex Heyne
Artocarpus integer (Thunb.) Merr.
Shorea venulosa Wood ex Meijer
Pouteria malaccensis (Clarke) Baehni
Dead
Shorea rubra P.S Ashton
Shorea maxwelliana King
112 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Sindora leiocarpa Backer ex de Wit
Palaquium cochlearifolium van Royen
Myristica beccarii Warb.
Hydnocarpus kunsteri (King) Warb.
Shorea maxwelliana King
Dead
Pouteria malaccensis (Clarke) Baehni.
Shorea maxwelliana King
Dead
Shorea parvifolia Dyer
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Gluta wallichii (Hk.f) Ding Hou
Shorea maxwelliana King
Shorea parvifolia Dyer
Hydnocarpus kunstleri (King) Warb.
Dead
Shorea maxwelliana King
Dead
Syzygium sp.2
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Shorea macroptera Dyer
Lithocarpus cantleyanus (King & Hook.f) Rehd
Gymnacranthera contracta Warb.
Lithocarpus cantleyanus (King & Hook.f) Rehd
Cleistanthus ferrugineus M.A.
Shorea beccariana Burck
Litocarpus cantleyanus (King & Hook.f) Rehd
Dipterocarpus lowii Hook.f
Shorea maxwelliana King
Swintonia glauca Engl.
Swintonia glauca Engl.
Shorea maxwelliana King
Hydnocarpus kunsteri (King) Warb.
A Field Guide to the Permanent Sample Plots • 113
Table 10. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
Species
Aporusa antennifera Airy Shaw
Shorea maxwelliana King
Koompassia malaccensis Maing. ex Benth.
Shorea venulosa Wood ex Meijer
Baccauera sumatrana M.A
Chionanthus pubicalyx (Ridl.) Kiew.
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Shorea venulosa Wood ex Meijer
Nephelium maingayi Hiern.
Myristica beccarii Warb.
Dead
Cleistanthus bakonensis Airy shaw
Shorea exelliptica Meijer
Agtathis borneensis Warb.
Dipterocarpus stellatus Vesque
Calophyllum lowii Hk.f.
Shorea exelliptica Meijer
Madhuca erythrophylla (K.ex G.) H.J.Lam
Shorea venulosa Wood ex Meijer
Dipterocarpus lowii Hook.f
Gonystylus forbesii Gilg.
Calophyllum lowii Hook.f.
Dead
Shorea maxwelliana King
Myristica beccarii Warb.
Cleistanthus ferrugineus M.A.
Shorea maxwelliana King
Nephelium maingayi Hiern.
100
80
60
40
20
0
0
0
59
11
227
58
214
55
203
18
1
2
39
61
20
3
281
4
40
208
215
105
36
107
104
110
97
223
135
134
23
278
168
245
216
170 136
213
224 137
20
40
54
40
199
131
112
5
92 91
88
113
268
93
90
114
16
17 193
30
128
86
127
60
87
207
151
60
44
253
6
287
81
267
204
80
52
202
286
80
80
222
120
161
8
13 123
160
155
257 49
212 45
12
152
153
7
173
32
229
35
77
14
198
73
289
166
75 247
246
124
169
100
126
284
100
100
80
60
40
20
0
Figure 13. Distribution of Trees in Plot RIL11
Lauraceae
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Dipterocarpaceae
Clusiaceae
Arecaceae
Verbenaceae
Theaceae
Sapotaceae
Sapindaceae
Rosaceae
Rhizophoraceae
Podocarpaceae
Oleaceae
Myrtaceae
Meliaceae
FAMILY
Anacardiaceae
Linaceae
RIL11
114 • A Field Guide to the Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 115
Table 11. List of Trees in Plot RIL11
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Baccaurea sumatrana M.A.
Syzygium bankensis (Hassk) Merr. & Perry
Shorea venulosa Wood ex Meijer
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Shorea parvifolia Dyer
Syzygium attenuatum Merr & Perry
Shorea venulosa Wood ex Meijer
Calophyllum lowii Hk.f.
Pimeleodendron papaveroides J.J.S.
Gluta wallichii (Hk.f) Ding Hou
Gluta wallichii (Hk.f) Ding Hou
Dipterocarpus lowii Hook.f.
Shorea exelliptica Meijer
Gluta wallichii (Hk.f) Ding Hou
Shorea maxwelliana King
Gluta wallichii (Hk.f) Ding Hou
Tetramerista glabra Miq.
Calophyllum lowii Hk.f
Shorea parvifolia Dyer
Dead
Dead
Gluta wallichii (Hk.f) Ding Hou
Dipterocarpus lowii Hook.f
Baccaurea sumatrana M.A
Dipterocarpus stellatus Vesque
Ptenandra galeata Cogn
Tetramerista glabra Miq.
Palaquium cochlearifolium van Royen
Swintonia glauca Engl.
Litsea nidularis Gamle
Teijsmaniodendron simplicifolium Merr.
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Shorea rubra Ashton
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus lowii Hook.f
116 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Shorea venulosa Wood ex Meijer
Eugeissona utilis Becc.
Eugeissona utilis Becc.
Syzygium sp.
Santiria laevigata Bl.
Palaquium cochlearifolium van Royen
Gluta wallichi (Hk.f ) Ding Hou
Palaquium cochlearifolium van Royen
Palaquium cochlearifolium van Royen
Syzygium sp.
Drypetes kikir Airy Shaw
Shorea rubra Ashton
Palaquium cochlearifolium van Royen
Shorea parvifolia Dyer
Carallia brachyata (Lour.) Merr.
Palaquium cochlearifolium van Royen
Shorea maxwelliana King
Koompassia malaccensis Maing. ex Benth.
Dead
Dead
Hydocarpus kunstleri (King) Warb.
Syzygium bankensis (Haask) Merr. & Perry
Dipterocarpus lowii Hook.f
Palaquium cochlearifolium van Royen
Myristica beccarii Warb.
Baccaurea sumatrana M.A.
Palaquium cochlearifolium van Royen
Shorea beccariana Burck.
Myristica beccarii Warb.
Shorea maxwelliana King
Myristica beccarri Warb.
Swintonia glauca Engl.
Shorea parviifolia Dyer
Dipterocarpus lowii Hook.f
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Elaeocarpus longipetiolatus Merr.
A Field Guide to the Permanent Sample Plots • 117
Table 11. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Shorea rubra Ashton
Syzygium clhoranthum (Duthie ) Merr.& Perry
Koompassia malaccensis Maing ex Benth.
Dead
Dipterocarpus stellatus Vesque
Palaquium cochlearifolium van Royen
Ctenolophon pavifolium van Royen
Ctenolophon parvifolium van Royen
Calophyllum pulcherrimum Wall.ex Choisy
Dipterocarpus stellatus Vesque
Diospyros convertifolra (Hiern) Bakh.
Shorea venulosa Wood.ex Meijer
Shorea maxwelliana King
Eugeissona utilitis Becc.
Unidentified
Artocarpus odoratissima Blanco
Shorea agami Ashton
Shorea parvifolia Dyer
Lithocarpus cantleyanus (King Hk.f) Rehd.
Shorea parifolia Dyer
Shorea agami Ashton
Syzygium chlorantum (Duthie) Merr. & Perry
Palaquium ferox H.J.Lam
Litsea machilifolia Gamble
Amoora lanceolata Hiem.
Tetramerista glabra Miq.
Calophyllum lowii Hk.f
Lithocarpus cantleyanus (King & Hk.f) Rehd.
Gluta wallichii (Hk.f) Ding Hou
Tetramerista glabra Miq.
Shorea macroptera Dyer
Shorea maxwelliana King
Shorea beccariana Burck
Lithocarpus cantleyanus (King Hk.f) Rehd
Crypteronia grifitii Clarke
Shorea beccariana Burck
Gluta wallichii (Hk.f) Ding Hou
118 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea venulosa Wood & Meijer
Aporusa antennifera Airy Shaw
Chionanthus ellipticus Bl.
Scaphium macropodum (Miq.) Beumée ex Heyne
Scaphium macropodum (Miq.) Beumée ex Heyne
Licania spendens (Korth) Prance
Pholidocarpus sp
Paratocarpus forbesii King
Sysygium chloranthum (Duthie) Mer. & Perry
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Shorea macroptera Dyer
Diospyros dayakensis Bakh
Dipterocarpus stellatus Vesque
Vatica umbonata (Hook.f.) Burck
Shorea venulosa Wood ex Meijer
Dipterocarpus lowii Hook.f.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Teijsmaniodendron simplicifolium Merr.
Shorea beccariana Burck
Shorea macroptera Dyer
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Santiria laevigata Bl.
Pternandra galeata Cogn.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Dead
Shorea beccariana Burck
Syzygium bankensis (Hassk.) Merr & Perry
Madhuca erytrophylla (K. ex G.) H. J. Lam
Podocarpus neriifolius D.Don
Dipterocarpus stellatus Vesque
Syzygium chloranthum (Duthie) M. & P.
A Field Guide to the Permanent Sample Plots • 119
Table 11. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea maxwelliana King
Hydnocarpus kunstleri (King) Warb.
Shorea beccariana Burck
Shorea maxwelliana King
Shorea rubra Ashton
Dead
Palaquium cochlearifolium van Royen
Calophyllum lowii Hk.f.
Shorea parvifolia Dyer
Lithocarpus cantleyanus (King &Hk.f.) Rehd.
Shorea pinanga Schreff.
Shorea pinanga Schreff.
Shorea rubra Ashton
Ternstroemia aneura Miq.
Dipterocarpus stellatus Vesque
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Calophyllum lowii Hk.f.
Syzygium bankensis (Hassk) Merr & Perry
Gluta wallichii (Hk.f.) Diong Hou
Lithocarpus catleyanus (King & Hk.f.) Rehd.
Amoora lanceolata Hiern.
Garcinia maingayi Hook.f.
Elaeocarpus longipetiolatus Merr.
Dead
Shorea beccariana Burck
Shorea rubra Ashton
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Swintonia glauca Engl.
Dead
Dipterocarpus stellatus Vesque
Shorea venulosa Wood ex Meijer
Lithocarpus cantleyanus (King &Hk.f.) Rehd.
Dead
Agathis borneensis Warb.
Sindora leiocarpa Backer ex de Wit.
Pternandra galeata D. Don
Shorea maxwelliana King
120 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Swintonia glauca Engl.
Baccaurea kunstleri King
Calophyllum lowii Hk.f.
Dipterocarpus stellatus Vesque
Sarcotheca diversifolia (Miq.) Hall.f.
Xanthophyllum vitellinum Bl.
Dacryodes rugosa (Bl.) H.J.Lam
Shorea venulosa Wood ex Meijer
Buchanania arborescens (Bl.) Bl.
Tetramerista glabra Miq.
Nephelium lappaceum Miq.
Elaeocarpus longipetiolatus Merr.
Syzygium bankensis (Hassk.) Merr. & Perry
Calophyllum pulcherrimum Wall. ex Choisy.
Cryptocarya tomentosa Bl
Xylopia altissima Boerl.
Lithocarpus cantleyanus (King & Hk.f.) Rehd
Elaeocarpus longipetiolatus Merr.
Shorea venulosa Wood ex Meijer
Dipterocarpus stellatus Vesque
Palaquium cochlearifolium van Royen
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (Kinf & Hk.f.) Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Agathis borneensis Warb
Shorea beccariana Burck
Shorea macroptera Dyer
Shorea venulosa Wood ex Meijer
Syzygium bankensis (Hassk.) Merr. & Perry
Shorea maxwelliana King
Syzygium bankensis (Hassk.) Merr. & Perry
Agathis borneensis Warb.
Shorea macroptera Dyer
Shorea maxwelliana King
Artocarpus anisophyllus Miq.
Amoora lanceolata Hiern.
A Field Guide to the Permanent Sample Plots • 121
Table 11. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea beccariana Burck
Aporusa prainiana King ex Gage
Dialium maingayi Baker
Gymnacranthera contracta Warb.
Shorea parvifolia Dyer
Madhuca erythrophylla (K. ex G.) H.J.Lam
Podocarpus neriifolius D. Don
Palaquium cochlearifoium van Royen
Shorea maxwelliana King
Shorea venulosa Wood ex Meijer
Gymnacranthera contracta Warb.
Shorea parvifolia Dyer
Myristica beccarii Warb.
Dead
Koompassia malaccensis Maing. ex Benth.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Shorea beccariana Burck
Atuna excelsa (Jack) Kosterm.
Heritiera javanica (Bl.) Kosterm.
Prunus arborea (Bl.) Kalkm.
Koompassia malaccensis Maing ex Benth.
Litsea nidularis Gamble
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Ternstroemia aneura Miq.
Pternandra galeata Cogn.
Dead
Syzygium chloranthum (Duthie) M. & P.
Syzygium chloranthum (Duthie) M. & P.
Lithocarpus catleyanus (King & Hk.f.) Rehd.
Dead
Syzygium caudatilimbum Merr.& Perry
Shorea parvifolia Dyer
Ctenolophan parifolius Oliver
Canarium littorale Bl.
122 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Lithocarpus cantleyanus (King Hk.f) Rehd.
Shorea parvifolia Dyer
Shorea maxwelliana King
Dead
Artocarpus integer (Thunb) Merr.
Gluta wallichii (Hk.f) Ding Hou
Shorea beccariana Burck
Gluta wallichii (Hk.f) Ding Hou
Gluta wallichii (Hk.f) Ding Hou
Dyera costulata (Miq) Hk.f
Shorea parvifolia Dyer
Dialium maingayi Baker
Dyera costulata (Miq) Hk.f
Dead
Dead
Calophyllum pullcherrimum Wall ex Planch.
Dipterocarpus lowii Hook.f
Dead
Dipterocarpus lowii Hook.f
Anisoptera megistocarpa Slooten.
Lithocarpus cantleyanus (King Hk.f) Rehd
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
Dead
Nephelium maingayi Hiern.
Shorea agami Ashton
Shorea parvifolia Dyer
Garcinia bancana Miq.
Nephelium maingayi Hiern.
Tetramerista glabra Miq.
Cleistanthus bakonensis Airy Shaw
Pholidocarpus sp.
Dead
Prunus arborea (Bl.) Kalkm.
Unidentified
Unidentified
Unidentified
100
80
60
40
20
0
215
223
0
1
2
5
6
84
85
92
94
93
95
104
212
9
7
97 98
102
101
211
214
213
203
205
204
105
8
89
88
90
107 106
99
20
219
218
3 4
87
20
108
209 207
208
210
217
220 221 226 227
222
225
224
216
96
103
100
0
199
82
116
111
109 110
117
195
12
118
13
11
14
40
15
80
79
19
124
122
121
120
119
188
77
16
17
66
76
245
135
18
20
60
140
60
24
22
70
71
25
29
58
30
33
31
27
28
59
62
63
139
80
60
61
177
178
167
168
39
34
35 36
44
54
149
165
166
164
163
38
37
53
57
143
147
162
56
150
42
41 40
45
48
49
154
152
100
43
46
47
52
55
146
145
51
156
155 157
148
285
100
286
274
284
158 161
159
270
283 273
280
281
282
276
277
279
269
265
80
266
267
169
268
179
175
137 138
141
142
26
181
171
180
260
262 261
263
259
264
172 173 174
257
256
128 136
134
23 21
69 68
67
73
74
72
127
133
187
186
184
183 182
191
185
75 126
125
130
244
192
243
193
242
194
132
131
189
241
252
251
249 253 254
250
246
247
232
40
190
114
115 202
200
237
240
201
231
234
235
239
238
229
230
113 112
198
197
236
233
228
100
80
60
40
20
0
Figure 14. Distribution of Trees in Plot RIL12
Flacourtiaceae
Fagaceae
Fabaceae
Euphorbiacea
Elaeocarpaceae
Ebenaceae
Dipterocarpaceae
Crypteroniaceae
Clusiaceae
Celastraceae
Burseraceae
Bombacaceae
Arecaceae
Araucariaceae
Apocynaceae
Annonaceae
Verbenaceae
Ulmaceae
Theaceae
Sterculiaceae
Sapotaceae
Sapindaceae
Rubiaceae
Rosaceae
Rhizophoraceae
Polygalaceae
Podocarpaceae
Oxalidaceae
Oleaceae
Myrtaceae
Myristicaceae
Meliaceae
Melastomataceae
Lecythidaceae
FAMILY
Anacardiaceae
Lauraceae
RIL12
A Field Guide to the Permanent Sample Plots • 123
124 • A Field Guide to the Permanent Sample Plots
Table 12. List of Trees in Plot RIL12
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea exelliptica Meijer
Artocarpus kemando Miq.
Shorea beccariana Burck
Pternandra galeata Cogn.
Canarium littorale Bl.
Shorea brunnescens Ashton
Pternandra galeata Cogn.
Syzygium ochneocarpa Merr.
Chionathus ellipticus Bl.
Pternandra galeata Cogn.
Diospyros oblongifolia Wall. ex G. Don
Baccaurea pendula Merr.
Pometia pinnata Forst.
Shorea parvifolia Dyer
Syzygium sp.
Dysoxylum alliaceum Bl.
Calophyllum lowii Hk.f.
Shorea ovalis (Korth.) Bl.
Koompassia malaccensis Maing. ex Benth.
Syzygium sp.
Carallia brachiata (Lour.) Merr.
Sindora velutina Baker
Lithocarpus pulcher (King) Markgr.
Oncosperma horridum Scheff.
Shorea maxwelliana King
Dipterocarpus lowii Hook.f.
Elaeocarpus sp.
Litsea nidularis Gamble
Shorea ovalis (Korth.) Bl.
Shorea maxwelliana King
Calophyllum lowii Hk.f.
Dead
Ternstroemia coriacea Sheff.
Shorea brunnescens P.S. Ashton
Sindora leiocarpa Backer ex de Wit
Syzygium grandis Wight.
Gluta wallichii (Hk.f.) Ding Hou
A Field Guide to the Permanent Sample Plots • 125
Table 12. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dipterocarpus stellatus Vesque
Shorea exelliptica Meijer
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Teijsmaniodendron simplicifolium Merr.
Dyera costulata (Miq.) Hk.f.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Artocarpus integer (Thunb.) Merr.
Shorea parvifolia Dyer
Pternandra galeata Cogn.
Syzygium albidiramea Merr.
Podocarpus neriifolius D. Don
Dead
Paratocarpus forbesii King
Dialium maingayi Baker
Syzygium chloranthum (Duthie) M.& P.
Santiria laevigata Bl.
Agathis borneensis Warb.
Alseodaphne ceratoxylon Kosterm.
Dialium maingayi Baker
Licania splendens (Korth.) Prance
Teijsmaniodendron simplicifolium Merr.
Dyera costulata (Miq.) Hk.f.
Hydnocarpus kunstleri (King) Warb.
Gymnacranthera contracta Warb.
Mezzetia parvifolia Becc.
Dead
Dead
Shorea ovalis (Korth.) Bl.
Koompassia malaccensis Maing. ex Benth
Baccaurea pendula Merr.
Syzygium acutangulum K.Schum.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Shorea parvifolia Dyer
126 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Sindora leiocarpa Backer ex de Wit
Dacryodes rugosa (Bl.) H.J.Lam
Shorea macroptera Dyer
Dead
Calophyllum lowii Hk.f.
Koompassia malaccensis Maing. ex Benth.
Dead
Lithocarpus bennetii (Miq.) Rehd.
Dead
Syzygium ochneocarpa Merr.
Lophopetalum subobovatum King
Pternandra caerulescens Jack.
Shorea ovalis (Korth.) Bl.
Gluta wallichii (Hk.f.) Ding Hou
Syzygium chloranthum (Duthie) M.& P.
Tetramerista glabra Miq.
Unidentified
Shorea pinanga Scheff.
Myristica villosa Warb.
Tetramerista glabra Miq.
Palaquium rostratum (Miq.) Burck
Cleistanthus myrianthus Krz.
Memecylon myrsinoides Blume
Dipterocarpus lowii Hook.f.
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Vatica sp. 1
Shorea macroptera Dyer
Dipterocarpus stellatus Vesque
Vatica sp. 1
Shorea maxwelliana King
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Syzygium grandis Wight.
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Dialium maingayi Baker
A Field Guide to the Permanent Sample Plots • 127
Table 12. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Dipterocarpus lowii Hook.f.
Teijsmaniodendron simplicifolium Merr.
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus lowii Hook.f.
Hopea sp.
Gluta wallichii (Hk.f.) Ding Hou
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus stellatus Vesque
Agathis borneensis Warb.
Shorea macroptera Dyer
Diospyros confertiflora (Hiern.) Bakh.
Dead
Dipterocarpus stellatus Vesque
Lithocarpus blumeanus (Korth.) Rehd.
Endiandra sp.
Licania splensens (Korth.) Prance
Dipterocarpus lowii Hook.f.
Dead
Agathis borneensis Warb
Teijsmaniodendrum simplicifolium Merr.
Shorea exelliptica Meijer
Agathis borneensis Warb.
Dipterocarpus stellatus Vesque.
Dipterocarpus stellatus Vesque.
Memecylon sp.
Gluta wallichii (Hk.f.) Ding Hou
Artocarpus anisophyllus Miq.
Shorea exelliptica Meijer
Pternandra galeata Cogn.
Shorea parvifolia Dyer
Mesua congestiflora P.F. Stevens
Agathis borneensis Warb.
Dead
Dialium maingayi Baker
Santiria griffithii (Hk.f.) Engl.
Shorea maxwelliana King
Shorea maxwelliana King
128 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Swintonia glauca Engl.
Dipterocarpus lowii Hook.f.
Dead
Xanthophyllum vitellinum (Bl.) Dietr.
Dead
Durio lanceolatus Mast.
Scaphium macropodum (Miq.) Beumée ex Heyne
Licania splendens (Korth.) Prance
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Shorea maxwelliana King
Lithocarpus cantleyanus (King & Hk.f.) Rehd
Amoora lenceolata Heim
Elaeocarpus longipetiolatus Merr.
Madhuca palembanica (Miq.) P.C.Yii & P.Chai
Hydnocarpus kunstleri (King.) Warb.
Shorea beccariana Burck
Lithocarpus bennetii (Miq.) Rehd.
Koompassia malaccensis Maing. ex Benth
Dipterocarpus stellatus Vesque.
Shorea beccariana Burck
Dead
Xanthophyllum amoenum Chod.
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Hidnocarpus kunstleri (King) Warb.
Hidnocarpus kunstleri (King) Warb.
NONE
Shorea rubra Ashton
Shorea ochracea Sym.
Shorea maxwelliana King
Dialium maingayi Baker
Vatica umbonata (Hook.f.) Burck.
Palaquium calophyllum (T.ex B.) Pierre
Shorea exelliptica Meijer
Vatica albiramis Sloot.
Aglaia ganggoo Miq.
A Field Guide to the Permanent Sample Plots • 129
Table 12. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Shorea pauciflora King
Agthis borneensis Warb.
Lithocarpus lucidus (Roxb.) Rehd.
Paratocarpus forbesii King
Gluta wallichii (Hk.f) Ding Hou
Shorea pauciflora King
Gymnacranthera contracta Warb.
Sarcotherca diversifolia (Miq.) Hall.f
Tetramerista glabra Miq.
Nauclea calysina (Bartl.) Merr.
Dead
Dipterocarpus lowii Hook.f
Koompassia malaccensis Maing. ex Benth.
Dipterocarpus lowii Hook.f
Sageraea elliptica (DC.) Hook.f.ex Th.
Diospyros everettii Merr.
Shorea brunnescens P.S.Ashton
Pometia pinnata Forst.
Santinia laevigata Bl.
Dacryodes rugosa (Bl.) H.J.Lam
Dead
Gluta wallichii (Hk.f.) Ding Hou
Madhuca magnifica H.J.Lam
Baccaurea edulis Merr.
Durio testudinarum Becc.
Dialium maingayi Baker
Xanthophyllum amoenum Chod.
Syzygium grandis Wight.
Shorea ovalis (Korth.) Bl.
Barinngtonia scortechinii King
Dyera costulata (Miq.) Hk.f.
Podocarpus neriifolius D.Don
Mezzetia parviflora Becc.
Horsfieldia crassifolia (Hk.f. et Th.) Warb.
Gymnacranthera contracta Warb.
Shorea atrinervosa Sym.
Shorea macroptera Dyer
130 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea pauciflora King
Teijsmaniodendron simplicifolium Merr.
Dipterocarpus lowii Hook.f.
Crypteronia sp.
Calophyllum depresinervosa Hend. &Wyatt-Sm.
Castanopsis motleyana King
Syzygium tawahense (Korth.) Merr.
Alseodaphne ceratoxylon Kosterm.
Diospyros laevigata Bakh.
Mezzetia parviflora Becc.
Shorea parvifolia Dyer
Barringtonia scortechinii King
Shorea macroptera Dyer
Tetramerista glabra Miq.
Shorea ochracea Sym.
Artocarpus odoratissima Blanco
Beilschmiedia sp.
Swintonia glauca Engl.
Memecylon sp.
Syzygium sp.
Syzygium sp.
Myristica beccarii Warb.
Xanthophyllym parvum Chod.
Pternandra galeata Cogn.
Licania splendens (Korth.) Prance.
Shorea parvifolia Dyer
Lithopcarpus cf. pseudokunstleri A. Camus
Myrisatica iners Bl.
Artocarpus integer (Thunb.) Merr.
Vatica umbonata (Hook.f.) Burck.
Hydnocarpus kunstleri (King) Warb.
Teijsmaniodendron simplicifolium Merr.
Dead
Lithocarpus ewyckii (Korth.) Rehd.
Amoora lanceolata Hiern.
Dead
Lithocarpus ewyckii (Korth.) Rehd.
A Field Guide to the Permanent Sample Plots • 131
Table 12. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
Species
Baccaurea pendula Merr.
Heritiera simplicifolia (Mast.) Kostrm.
Swintonia glauca Engl.
Baccaurea sumatrana M. A.
Garcinia amingayi Hook.f.
Gluta wallichii (Hk.f.) Ding Hou
Licania splendens (Korth.) Prance
Swintonia glauca Engl.
Hydnocarpus kunstleri (King) Warb.
Celtis philippensis Blanco
Shorea maxwelliana King
Dead
Dead
Swintonia glauca Engl.
Shorea maxwelliana King
Lithocarpus cantleyanus (King & Hk.f.) Rehd.
Agathis borneensis Warb.
Shorea rubra Ashton
Aporusa antennifera Airy Shaw
Memecylon myrsinoides Blume
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea maxwelliana King
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Shorea brunnescens P.S. Ashton
Amoora lanceolata Hiem
Dipterocvarpus stellatus Vesque
The Center for International Forestry Research (CIFOR) is a leading international
forestry research organization established in 1993 in response to global concerns about the
social, environmental, and economic consequences of forest loss and degradation. CIFOR
is dedicated to developing policies and technologies for sustainable use and management
of forests, and for enhancing the well-being of people in developing countries who rely on
tropical forests for their livelihoods. CIFOR is one of the 15 Future Harvest centres of the
Consultative Group on International Agricultural Research (CGIAR). With headquarters
in Bogor, Indonesia, CIFOR has regional offices in Brazil, Burkina Faso, Cameroon and
Zimbabwe, and it works in over 30 other countries around the world.
Donors
The Center for International Forestry Research (CIFOR) receives its major funding from
governments, international development organizations, private foundations and regional
organizations. In 2005, CIFOR received financial support from Australia,Asian Development
Bank (ADB), Belgium, Brazil, Canada, China, Centre de coopération internationale
en recherche agronomique pour le développement (CIRAD), Cordaid, Conservation
International Foundation (CIF), European Commission, Finland, Food and Agriculture
Organization of the United Nations (FAO), Ford Foundation, France, German Agency for
Technical Cooperation (GTZ), German Federal Ministry for Economic Cooperation and
Development (BMZ), Indonesia, International Development Research Centre (IDRC),
International Fund for Agricultural Development (IFAD), International Tropical Timber
Organization (ITTO), Israel, Italy, The World Conservation Union (IUCN), Japan, Korea,
Netherlands, Norway, Netherlands Development Organization, Overseas Development
Institute (ODI), Peruvian Secretariat for International Cooperation (RSCI), Philippines,
Spain, Sweden, Swedish University of Agricultural Sciences (SLU), Switzerland, Swiss Agency
for the Environment, Forests and Landscape, The Overbrook Foundation, The Nature
Conservancy (TNC), Tropical Forest Foundation, Tropenbos International, United States,
United Kingdom, United Nations Environment Programme (UNEP), World Bank, World
Resources Institute (WRI) and World Wildlife Fund (WWF).
The CIFOR Permanent Sample Plots (PSP) were
established in 1998 in conjunction with the Phase I of the
ITTO Project dealing with sustainable forest management,
integrated land use, reduced-impact logging, biodiversity
in the production forest, and cultural and social aspects
of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes,
long term growth and yield and to provide critical data for
evaluation of ecological models. For silvicultural purposes
PSP supply data on diameter and volume increment as
well as stand structure dynamics.
In particular, the PSP in the Malinau Research Forest
provides a valuable insight into the diversity of trees found
there and the growth of the forest. This guidebook has
been designed to facilitate the growing interests and needs
of visitors to the Malinau Research Forest (MRF).
PATEN MA LINAU
KAB U
I N TI M U N G