Human Evolution and Migration in a Variable Environment:
The Amplifier Lakes of East Africa
Martin H. Trauth',', Mark A. Maslin", Andreas G.N. Bergner', Alan Deino-, Annett Junginger=', Eric Odada",
Daniel O. Olago", Lydia Olaka=', Manfred R. Strecker'
'Institut fur Geowissenschaften,
Universitat Potsdam, Potsdam, Germany
bGeography Department, University College London, London, UK
cBerkeley Geochronology Center, 245 Ridge Road, Berkeley, California 94709, USA
dDFG Graduateschool "Shaping the Earth's Surface in a Variable Environment", University
<Department of Geology, University of Nairobi, Nairobi, Kenya
of Potsdam
*corresponding author at Institut fur Geowissenschaften,
Universitat Potsdam, Karl-Liebknecht-Str.
Tel. +49-331-977-5810,
Fax +49-331-977-5700,
E-mail [email protected]
24, 14476 Potsdam,
Germany.
The development
of the Cenozoic East African Rift System (EARS) profoundly
re-shaped
the
landscape and significantly increased the amplitude of short-term environmental
response to climate
variation. In particular, the development of amplifier lakes in rift basins after three million years ago
significantly contributed to the exceptional sensitivity of East Africa to climate change compared to
elsewhere on the African continent. These amplifier lakes respond rapidly to moderate, precessionalforced climate shifts, and as they so apply dramatic environmental
pressure to the biosphere. Rift
basins, when either extremely dry or lake-filled, form important barriers for migration, mixing and
competition of different populations of animals and hominins. Amplifier lakes link long-term, highamplitude tectonic processes and short-term environmental
fluctuations. East Africa may have been
the cradle of humankind as a consequence of this strong link between different time-scales.
The established drivers of environmental instability in East Africa are volcano-tectonic
phenomena
associated with the East African Rift System (EARS) and associated plateaus (time scales of 106 to 105
years), orbitally-driven changes in insolation (l05 to 104 years), and variations in total solar irradiance (103 to
10° years) (Kutzbach and Street-Perrott, 1985; Nicholson, 1996; Verschuren et al., 2000; Trauth et al.. 2003,
2005; Barker et aI., 2004; Sepulchre et al., 2006). In addition to these direct influences, remote drivers of
East African environmental changes include the uplift of the Tibetian plateau and the establishment of the
African-Asian monsoon system (106 to 105 years), orbitally-driven glacial-interglacial cycles (104 to 103
years), and sea-surface temperature variations driven by ocean-atmosphere phenomena such as the Indian
Ocean Dipole (lOD) and the EI Nino/Southern Oscillation (ENSO) (104 to 10° years) (e.g., Kutzbach and
Street-Perrott, 1985; Saji et al., 1999; Schreck and Sernazzi, 2004).
Drivers of environmental changes acting on relatively long time scales have the greatest largest amplitudes
and expectedly the greatest effect on the habitats of animals and hominins. For example, growth of the East
African and Ethiopian plateaus are considered to be major influences in the dramatic vegetation change that
has taken place on the continent during the last twenty million years (Sepulchre et al., 2006). The classic
savanna hypothesis of Henry Fairfield Osborn and Raymond Dart attempted to link the evolutionary
divergence of hominins and other great apes, and the emergence of bipedalism, with the forest-savanna
transition in Mio-Pliocene time (e.g., Dart, 1953). Though largely disproved, the savanna hypothesis was an
early attempt to link tectonic displacements, environmental change and hominin evolution.
.
The effects of such long-term tectonic displacements, however, can also have an influence on the magnitude
of short-term environmental instabilities, thus leading to a more direct impact on the biosphere. Ancient
shorelines of East African lakes, most of them reduced to salty puddles today, document orbitally-driven
lake-level fluctuations of tens or hundreds of meters as a response to only moderate climate shifts (e.g.,
Washbourn-Kamau, 1970, 1975; Street-Perrott and Harrison; 1985; Trauth et al., 2003; Bergner et al., 2003;
Garcin et al., 2009) (Fig. 1). This is related to the amplification effect, the result of a tectonically-formed
graben morphology in combination with an extreme contrast between high precipitation in the elevated parts
of the catchment and high evaporation in the lake area (Street-Perrott and Harrison. 1985; Bergner et al., in
press).
The recent lake-level history of Lake Naivasha, and adjacent lakes in the Kenya Rift, highlights the extreme
sensitivity of such amplifier lakes to climate changes (Fi~~.2) (Washbourn, 1975; Hastenrath and Kutzbach,
1983; Bergner et al., 2003; Trauth et aI., 2003, 200S: USDAiNASA/RaytheonfUMD;
Ministry of Water and
Irrigation of Kenya, 2008) (see supplementary information). On the time scale of 10° to 101 years, Lake
Naivasha typically varies 0-1.5 m in lake level (USDAINASA!RaytheonIUMD;
Ministry of Water and
Irrigation of Kenya. 2008), whereas lake-level fluctuations on a time scale of 102 to 103 years are on the
order of tens of meters (Verschuren et al., 2000). Early Holocene water-level variations reach amplitudes of
up to one hundred meters (Washbourn, 1975: Hastenrath and Kutzbach, 19R3: Bergner et al., 20(3), whereas
the lake level varied by -150 m on a time scale of 104 to 105 years (Bergner ct al., 2003: Trauth ct al.. 2003).
Even greater water depths of more than 250 m are documented for a time scale of 105 to 106 years, after
about three million years ago (Trauth et a!., 2005, 2007).
Essentially, water-level changes in the Naivasha basin follow a power law where larger lake-level variations
occur on longer time scales (Fig. 2). Other African lakes such as Lake Chad (Leblanc et al., 2006) and Lake
Victoria (Hastenrath and Kutzbach, 1983) respond over similar time scales with significantly lower lakelevel variations, whereas some lakes, e.g., Lake Ziway (Gillespie et al., 1983), Lake Nakuru (Wash bournKamau, 1970) or Lake Suguta (Garcin et aI., 2009) demonstrate more extreme lake-level variations than
Lake Naivasha. Importantly, all amplifier lakes occur in the tectonically-controlled
environments of the
Kenyan and Ethiopian rifts (Fig. I and 2). Both the basin morphology and precipitation-evaporation
contrasts
in the catchment area of the amplifier lakes are the result from extensional tectonic processes. Thus, this
study highlights the relevance of long-term tectonic processes to short-term climate fluctuations and
environmental instability.
When did the amplifer lake systems evolve in East Africa? The volcano-tectonic segmentation of the Kenya
Rift into smaller, partly internally drained basins began during the Pliocene and continues into the present
day. Between 5.5 and 3.7 Ma, earlier halfgraben structures were antithetically faulted, thus creating a fullgraben morphology in most parts of the rift (Strecker et al., 1990; KRISP Working Party. 1991: Ebinger et
al., 1998: and supplementary information). The internal drainage conditions were caused by the conspiring
effects of magmatic activity centered along the volcano-tectonic axis and the formation of central volcanoes
during the Pleistocene. Importantly, these processes were augmented by the formation of horst and graben
structures that fundamentally influenced the fluvial network of the rift. The timing of these processes
coincides with a re-orientation of the extension direction from an earlier E-W to a neotectonic ESE-WNW
orientation during Plio-Pleistocene (Strecker et aI., 1990; and supplementary information).
The hallmark of these young extensional processes is the formation of presently isolated sub-basins that
coalesced multiple times in the past during episodes of climatic changes when East Africa must have
experienced greater amounts of precipitation. The combination of these tectonic movements and the
existence of earlier volcanic edifices results in the distinct precipitation-evaporation contrast between the rift
shoulders receiving more than 2,000 mm of mean-annual rainfall and graben areas with up to 4,000 mm of
evaporation today (Garcin et al., 20(9). Interestingly, the development of amplifier lake systems at that time
is also documented in a distinct change in the paleo-lake character. Whereas the -4.6-4.5 Ma old paleo-Lake
Turasha in the central Kenya Rift was a relatively stable, large and shallow freshwater lake as documented
by a -90 m thick sequence of diatomites in the Naivasha basin, the younger paleo-lakes Gicheru (-1.9-1.6
Ma) and Kariandusi (-1.0-0.9 Ma) were characterized by water depths of > 150 meters that varied
significantly on all time scales and may thus present the transition to the amplifier-lake conditions (Trauth et
al., 2005.2007).
The switch from extensive, possibly shallow rift lakes during the early halfgraben stages to amplifier lakes
Plio-Pleistocene time is important as it made this region very sensitive to climate changes. Sensitivity alone,
however, does not alter the local environment as an initial change in climate is required for a response to
occur. There is a growing body of geologic evidence for precession-forcing of moisture availability in the
tropics, both in East Africa and elsewhere during the Plio-Pleistocene (Bush et al., 2002; Deino et al., 2006;
Kingston et al., 2007; Hopley et aI., 2007; Trauth et al., 2003, 2007; Cruz et a!. 2005; Wang et (11., 2004:
Lepre et al., 2007). The precessional control on tropical moisture has also been clearly illustrated by climate
modeling of Clement et al. (2004), which showed that a 180· shift in precession could change the annual
precipitation in the tropics by at least 180 mm/year and induce a significant shift in seasonality, whereas
precession has almost no influence on global or regional temperatures (Clement et al., 20(4). A synopsis of
lake-level histories in the EARS suggests that precessional forcing of moisture availability fundamentally
influences the expansion and shrinkage of lakes. The first possible environmental state occurs at minimum
precession and maximum insolation in the Northern Hemisphere, inducing a wetter climate over East Africa
(Kutzbach and Street-Perrott, 1985).
r
Once a critical evaporation/precipitation
threshold is passed, the amplifier lakes respond rapidly (within less
than 200 years) and become very large and deep (Kutzbach and Street-Perrott, 1985; Bergner et al., 2003;
Garcin et aI., 2009). The presence of such large lakes affects the local environment by altering the adiabatic
lapse rates leading to generally conditions, including local vegetation changes and forest expansion.
Response times between climate and major vegetation changes can be as short as 50 years (Hughen et al.,
2004; Maslin, 2004). Figure 3a illustrates the effect of the growth of an amplifier lake and the expansion of
surrounding forest during this Wet Period. Once the maximum extent of rift lakes was established, on a
regional basis they would have served as an east-west barrier to the migration of animals and early hominins.
North-south movement between the different lake systems would have been relatively unimpeded, but with
rich fertile woody environments surrounding lakes and on the rift shoulders, there would have been little or
no environmental pressure to do so. As the precession cycle progresses and the insolation maximum moves
towards the Southern Hemisphere, moisture availability in EARS would be reduced. The lakes initially have
a muted response as their presence maintains a relative wet environment in the rift valley, until increased
aridity overcomes inertia and the lakes shrink.
Modeling results and field observations suggest that East African lakes require up 2,000 years to disappear
during a wet-to-dry climate transition (Bergner et al., 2003; Garcin et al., 2009). Figure 3b illustrates this
Transitional Period and the effect of reduced rainfall and lake shrinkage on the adjacent vegetation mosaic,
with forests retreat from lakeshore environments to the rift shoulders. This facilitates both north-south and
east-west migration of biota. As the insolation maximum reaches the Southern Hemisphere the rift valley
becomes extremely arid. Dust volume carried on the wind to both Mediterranean and Indian Ocean sites
increases (deMenocal, 2004; Larrasoafia et aI., 2003; Trauth et al., 2009). The effects on the environment are
illustrated in Figure 3c, with small refugia of forest surviving only in the rift shoulder areas or on isolated
volcanic edifices during the Dry Period. This effect has been seen in flightless bush crickets, which were
forced into shrinken forest refugia on the rift shoulders during dry intervals, evolving into separate species
(Voje et al., 2008). We postulate that a dry rift valley has acted as a major barrier, isolating animals and
hominins periodically during the Plio-Pleistocene, resulting in small populations trapped in higher altitude
refugia with the possibility of allopatric speciation.
Consequently, alternating wet and very dry conditions superposed on a a changing tectonically active
environment in the EARS may have resulted in the formation of barriers to animal and hominin migration,
both east-west and north-south. The very dry periods could have isolated populations in small refugia with
the possibility of allopatric speciation. The Transitional Periods provide the most opportunity for migration
(Fig. 3b). However, these periods are also most sensitive to extreme global climate influences, for example
millennial-scale Heinrich or Dansgaard-Oeschger events or extreme EI Nifio/Southern Oscillation (ENSO)
intervals, because "the water balance of the amplifier lakes will react strongly to small changes in moisture
availability. These conditions may provide the required forcing of environmental conditions that constitutes
the basis of the variability selection hypothesis (Potts, 1998).
We can also speculate on the relative durations of wet periods based on the timing of the precessional cycle.
Precession is sinusoidal, with intervals of little or no change in insolation, followed by intervals of a much
increased rate of changes. Sinusoidal forcing results in intervals of by ~8,000 years when relatively little
change in daily insolation occurs (Maslin et al., 2005), representing the Wet Period and Dry Period intervals
described above. These are followed by a rapid change of about 2,500 years which 60% of the total variation
in daily insolation and seasonality occurs. These intervals of rapid change correspond to the Transitional
Periods described above. Nevertheless, without the tectonically created topographic boundary conditions
these rapid and extreme changes associated with the amplifier lakes could not have occured in these local
environments.
Although highly hypothetical, the close relationship between tectonics, climate, topography and superposed
oscillating environmental changes on all time scales must have fundamentally influenced evolutionary
processes in this tropical environment. The East African amplifier lakes are a keystone in this scenario and
may help explain why East Africa became the place where early humans evolved. The further understand
Plio-Pleistocene climate change in the EARS, we support proposed initiatives for scientific drilling of
hominin-related sedimentary basins (e.g., Cohen, 2009).
References and Notes
Barker, P.A., Talbot, M.R., Street-Perrott, F.A., Marret, F., Scourse, J.D., Odada, E., 2004. Late Quaternary
climatic variability in intertropical Africa. In: Battarbee, R.W., Gasse, F., Stickley, C.E. (Ed.), Past
;
.
Climate Variability through Europe and Africa. Developments in Paleoenvironmental Research. Kluwer
Academic Publishers, Dordrecht, pp. 117-138.
Bergner, A.G.N., Trauth, M.H., Bookhagen, B., 2003. Magnitude of precipitation / evaporation changes in
the Naivasha Basin (Kenya) during the last 150 kyrs. Global and Planetary Change 36, 117-136.
Bergner, AG.N., Strecker, M.R., Trauth, M.H., Deino, A., Gasse, F, Blisniuk, P., Duhnforth, M, in press.
Tectonic versus climate influences on the evolution of the lakes in the Central Kenya Rift. Quaternary
Science Reviews.
Bush, M.H., Miller, M.C., De Oliveira, P.E., Colinvaux, P.A., 2002. Orbital forcing signal in sediments of
two Amazonian lakes. Journal of Paleolimnology 27, 341-352.
Clement, AC., Hall., A., Broccoli, AJ., 2004. The importance of precessional signals in the tropical climate.
Climate Dynamics 22, 327-341.
Cohen, A., 2009, Connecting Scientific Drilling and Human Evolution. EOS, 90, 121-122.
Cruz, FW., Bums, S.J., Karmann, 1., Sharp, W.D., Vuille, M., Cardoso, A.O., Ferrari, J.A., Silva Dias, P.L.,
Viana, 0., 2005. Insolation-driven changes in atmospheric circulation over the past I I 6,000 years in
subtropical Brazil. Nature 434, 63-66.
Dart, R., 1953. The predatory transition from ape to man. International Anthropological and Linguistic
Review 1,201-219.
Deino, A.L., Kingston, J.D., Glen, J.M., Edgar, R.K., Hill, A., 2006. Precessional forcing of lacustrine
sedimentation in the late Cenozoic Chemeron Basin, Central Kenya Rift, and calibration of the Gauss/
Matuyama boundary. Earth and Planetary Science Letters 247, 41-60.
deMenocal, P.B., 2004. African climate change and faunal evolution during the Pliocene-Pleistocene. Earth
and Planetary Science Letters 220, 3-24.
Garcin, Y., Junginger, A., Melnick, D., Olago, D.O., Strecker, M.R., Trauth, M.H., 2009. Late PleistoceneHolocene rise and collapse of Lake Suguta, northern Kenya Rift. Quaternary Science Reviews 28,
911-925.
Gasse, F, 1977. Evolution of Lake Abhe (Ethiopia and TFAI), from 70,000 b.p. Nature 265, 45-47.
Gasse, F, 1990. Tectonic and climatic controls on lake distribution and environments in Afar from Miocene
to present. In: Katz, BJ. (Ed.), Lacustrine Basin Exploration, Case Studies, and Modern Analogs. AAPG
Memoir, pp. 1-34.
Gillespie, R., Street-Perrott, FA., Switsur, R., 1983. Post-glacial arid episodes in Ethiopia have implications
for climate prediction. Nature 306, 680-683.
Hastenrath, S., Kutzbach, J.E., 1983. Paleoclimatic Estimates from Water and Energy Budgets of East
African Lakes. Quaternary Research 19, 131-153.
Hopley, P.J., Marshall, J.D., Weedon, G.P., Latham, A.G., Herries, A.1.R., Kuykendall, K.L., 2007. Orbital
forcing and the spread of C4 grasses in the late Neogene: stable isotope evidence from South African
speleothems. Journal of Human Evolution 53, 620-634.
Hughen, K.A., Eglinton, T.I., Xu, L., Makou, M., 2004. Abrupt Tropical Vegetation Response to Rapid
Climate Changes. Science 304, 1955-1959.
Kingston, J., Deino, A., Hill, A., 2007. Astronomically forced climate change in the Kenyan Rift Valley 2.72.55 Ma: implications for the evolution of early hominin ecosystems. Journal of Human Evolution 53,
487-503.
Kutzbach, J.E., Streetperrott, FA., 1985. Milkankovitcb forcing of fluctuations in the level of tropical lakes
from 18 to 0 kyr BP. Nature 317, 130-134.
Larrasoafia 1. c., R., AP., Rohling, Winklhofer, M., Wehausen, R., 2003. Three million years of monsoon
variability over the northern Sahara. Climate Dynamics 21, 689-698.
Leblanc, M., Favreau, G., Maley, J., Nazoumou, Y., Leduc, c., Stagnitti, F, van Oevelen, P., Delclaux, F,
Lemoalle, L, 2006. Reconstruction of Megalake Chad using Shuttle Radar Topographic Mission data.
Palaeogeography, Palaeoclimatology, Palaeoecology 239, 16-27.
Lepre, c.r., Quinn, R.L., Joordens, J.C.A., Swisher III, c.c., Feibel, C.S., 2007. Climatic forcing of
terrestrial environmental change during the emergence of H. erectus: evidence from tbe KBS Member of
the Koobi Fora Formation. Journal of Human Evolution 53, 504-514.
Maslin, 2004. Ecological versus climatic thresholds. Science 306, 2197-2198.
Maslin, M.A., Mabli, Y., Phillips 0., Cowling S., 2005. New views on an old forest: assessing the longevity,
resilience and future of tbe Amazon Rainforest. Transactions of the Institute of British Geographers 30,
390-401.
Nicholson, S.E., 1996. A review of climate dynamics and climate variability in Eastern Africa. In: T.c.
Johnson, E.O.O. (Ed.), The Limnology, Climatology and Paleoclimatology of the East African Lakes, The
International Decade for the East African Lakes (IDEAL). Gordon and Breach, Newark, NJ, pp. 25-56.
Owen, R.B., Barthelme, ].W., Renaut, R.W., Vincens, A., 1982. Palaeolimnology and archaeology of
Holocene deposits north-east of Lake Turkana, Kenya. Nature 298, 523-529.
Party, K.W., 1991. Large scale variations in lithospheric structure along and across the Kenya Rift. Nature
317,130-134.
Potts, R., 1998. Environmental hypotheses of hominin evolution. Yearbook of Physical Anthropology 41,
93-136.
Richardson, J.L., Dussinger, R.A, 1986. Paleolimnology of mid-elevation lakes in the Kenya Rift Valley.
Hydrobiologia 143, 167-174.
Saji, N.H., Goswami, B.N., Vinayachandran, P.N., Yamagata, T., 1999. A dipole mode in the tropical Indian
Ocean. Nature 401,360-363.
Schreck, c., Semazzi, F., 2004. Variability of the recent climate of eastern Africa. International Journal of
Climatology 24, 681-701.
Sepulchre, P., Ramstein, G., Fluteau, F, Schuster M., Tiercelin, J.J., Brunet, M., 2006. Tectonic Uplift and
Eastern Africa Aridification. Science 313, 1419-1423.
Strecker, M.R., Blisniuk, P.M., Eisbacher, G.H., 1990. Rotation of extension direction in the central Kenya
Rift. Geology 18, 299-302.
Street-Perrott, FA., Harrison, S.P., 1985. Lake Level and Climate Reconstructions. In: Hecht, A.D. (Ed.),
Paleoclimate Analysis and Modeling. John Wiley and Sons, New York, pp. 291-340.
Sturchio, N.C., Dunkley, P.N., Smith, M., 1993. Climate-driven variations in geothermal activity in the
northern Kenya rift valley. Nature 362, 233-234.
Trauth, M., Deino, A., Bergner, A., Strecker, M., 2003. East African climate change and orbital forcing
during the last 175 kyr BP. Earth and Planetary Science Letters 206, 297-3 I3.
Trauth, M., Larrasoafia, 1., Mudelsee, M., 2009. Trends, rhythms and events in Plio-Pleistocene African
climate. Quaternary Science Reviews 28, 399-411.
Trauth, M.H., Deino, A., Strecker, M.R., 200 I. Response of the East African climate to orbital forcing during
the last interglacial (130-117 ka) and the early last glacial (117-60 ka) Geology 29, 499-502.
Trauth, M.H., Maslin, M.A., Deino, A., Strecker, M.R., Bergner, A.G.N., Duhnforth, M., 2007. High- and
low-latitude forcing of Plio-Pleistocene East African climate and human evolution. Journal of Human
Evolution 53, 475-486.
Trauth, M.H., Dei no, A.L., Bergner, A.G.N., Strecker, M.R., 2003. East African climate change and orbital
forcing during the last 175 kyr BP. Earth and Planetary Science Letters 206, 297-313.
Trauth, M.H., Maslin, M.A., Deino, A., Strecker, M.R., 2005. Late Cenozoic moisture history of East Africa.
Science 309, 2051-2053.
Verschuren, D., Laird, K.R., Cumming, B.F., 2000. Rainfal1 and drought in equatorial east Africa during the
past 1,100 years. Nature 403, 410-413.
Voje, K., Hemp, C.; Flagstad, 0., Saetre, G., Stenseth, N., 2008. Climatic change as an engine for speciation
in flightless Orthoptera species inhabiting African mountains. Molecular Ecology 18,93-108.
Wang, X., Auler, A., Edwards, Cheng, H., Cristalli, Smart, Richards, Shen, 2004. Wet periods in northeastern
Brazil over the past 210 kyr linked to distant climate anomal ies. Nature 432, 740-743.
Washbourn-Kamau, C.K., 1970. Late Quaternary Chronology of the Nakuru-Elmenteita
Basin, Kenya.
Nature 226, 253-254.
Washbourn-Kamau, C.K., 1975. Late Quaternary Shorelines of Lake Naivasha, Kenya. Azania X, 77-92.
Acknowledgments
This project was funded by grants to M.T. and by the German Research Foundation (DFG) and a UCL
Graduate School grant to M.M. We also thank the German Academic Exchange Service (DAAD) and the
DFG Graduate School GRKI364 for their support of LO. We thank the Government of Kenya (Research
Penn its MOST 13/001l30C 59110, 59118 and 59/22), the Ministry of Water and Irrigation of Kenya and the
University of Nairobi for research permits and support. We thank S. Higgins, F Schrenk, R. Tiedemann, L.
Epp and K. Stoof and the participants of the EAGLE 2008 workshop at the University of Potsdam for
inspiring discussions. We would also like to thank H. Douglas-Dufresne and Pete Ilsley of WildFrontiers
Kenya, and Benjamin Simpson and Michael Watson of TropicAir Kenya for logistical support during the
helicopter expeditions to the Suguta Valley in 2007 and 2008.
f
Figure Caption
Figure 1 Map of East Africa showing topography, faults and lake basins (modified from Trauth et al., 2005).
Note the location of amplifier, intermediate and non-amplifier lakes. All amplifier lakes are located in a rift
environment and respond sensitively to relatively moderate climate variation.
Figure 2 (a) Lake-level change in the Naivasha basin in the Central Kenya Rift based on lake sediments and
paleo-shorelines; (Naiv I) observations (Ministry of Water and Irrigation of Kenya, 2008), (Naiv 2) satellite
altimetry (USDNNASAiRaytheoniUMD),
(Naiv 3) sediment characteristics (Verschuren et al., 2000), (Naiv
4) sediment characteristics and paleo-shorelines (Richardson & Dussinger, 1986), (Naiv 5) sediment
characteristics, authigenic mineral phases, diatom assemblages (Trauth et al., 200 1; 2003), (Naiv 6) sediment
characteristics, diatom assemblages (Trauth et al., 2005, 2007). (b) Compilation of lake-level variations in
Africa at time scales from 10° to 107 years based on lake sediments and paleo-shorelines; Lake Victoria
(Hastenrath and Kutzbach, 1983), Lake Chad (Leblanc et al., 2006), Lake Turkana (Owen et al., 1982;
Hastenrath and Kutzbach, 1983), Lake Naivasha (Washboum, 1975; Hastenrath and Kutzbach, 1983;
Bergner et al., 2003; Trauth et aI., 2003, 2005. 2007; USDAiNASA/RaytheoniUMD;
Ministry of Water and
Irrigation of Kenya, 2008), Lake Ziway (Gillespie et al., 1983), Lake Abhe (Gasse et al., 1977), Lake
Nakuru-Elmenteita (Washbourn-Kamau.
1970), Lake Rukwa (+200 m) (Kervyn et al., 2006), and Lake
Suguta (Garcin et al., 2(09).
Figure 3 Conceptional model of orbitally-induced climate variation, environmental change, and population
speciation and radiation in a rift environment. (a) During precessional minima, hence Northem Hemisphere
insolation maxima, large rift lakes form natural barriers that inhibit mixing of the two separate populations I
and 2 of animals and hominins. New species evolve independently in geographic isolation according to the
model of allopatric speciation. Due to favorable environmental conditions during a wetter climate,
population sizes increase, the larger populations split, and sub-populations migrate to other regions with
favorable conditions. (b) Near the precession inflection point, favorable but deteriorating conditions foster
migration of portions of the population along a corridor through the fading barrier. The populations I and 2
meet and compete during a gradually drying climate and limited availability of water and food. The
populations utilize the margins of lakes to migrate southwards following favorable conditions in course of
latitudinal climate shifts. (c) During the precession maximum, severe aridity creates a dry valley as a new
barrier for migration of animals and early hominins forming two new populations 1 and 2. The limited
availability of water and food results in a reduction of population size, and a tendency to migrate toward
uplands where favorable conditions persist. Isolation of sub-populations 2a and 2b fosters allopatric
speciation.
6
Areas> 1000 m
above sea level
Normal Faulr
•
@
•
o
Amplifier Lakes
Intermediate Lakes
Non-Amplifier Lakes
Other sites described
in the main
_____________
~ ~ ':. s_ t _0
Indian Ocean
42
Trauth et al., Figure 1
44
.!
(b)
(a)
1
300
Lake Naivasha
250
...•
200
"
~
KILOYEARS·\ ~ :~~~~~~
E
-
g!
(NA11I51
150
ctN
Q)
.....J
.2i
.5
rURI[S-KILOYEARS
1900±60m
'NArV4j
V'M"Q;i:'r.',u,~~
100
MomH:'
lNAIV3i
/,""'V1I ",:"7;~':~
\ ..:
(Nt.IV"
\.
1
.-'1
r
O=J ...
~,"\.;_","1
10°
~, "~
"n' :
"I
10 1
f'!ll
10'
10'
~
'
10'
Time before present (years)
Trauth et aI., Figure 2
I
i
!
.1
v;- 200
I!
i. ,
a;
i
g!
ill
1
i
i
III
j
!i
'I
I
,I
105
ll
~
,.j'\v (:
•. '
'[I,
I
I
li
I
Ii
u
II
Ii
,II
10"
/:'
s
//
150
:.;','
.
1
,-.Y
~
~....
",",
. -V--i-
,fii
:;t-"'.'
~~,:
e.
'.d<'q,"
~:
:
~eV
:~\Ci
~q;,
\e~
\~
'V
(l)
-'"
..J 100
'"
@,
( ",
ol
10°
,,";
,.' .. 0
i$~i~·;:::··.
50
10'
:0 ",'"
P
I
",. ,lID
<ZI ~
" ~~
-»:,.:
<l>
..J
'vti
:,f,£ .... q} ...
....,
I
~.
.'~
~~~-,..~
,~ .~,.•
r.
n'
" -;-.Y.",
o:,.{
Q;
I'
~.. .:
18f!7h!5'1l
50
______'
,r'" .f'!
:t~.Jp'
.:J""<'" / ' '
-J'
Jl
,
9
.v,~. /
250"
1
1900±180rn
{NAlV6)
-
.s
.;, ,
0,.
African Lakes
-In ',
__
100 KYRS·MYRS
.,
300,
in'
10'
10"
10'
Tlrne before present (years)
10'
10·
10'
(a) Wet Period during
Precession Minimum
(c) Dry Period during
Precession Maximum
(b) Transition Period near
Precession Inflection Point
.~
55$
0.02
Min!rnurn
Transition
200
c
.Q 0.01
(f)
u:
(j)
0
o
r
150 ~
<1>
100 ~
(!)
G::
<
-0.01
50
-0.02
0
20
15
10
5
Time (103 Years)
Trauth et aI., Figure 3
0
~
:3
0.021
150
.~ 0.01
(/)
~
o
100 r:
(l)
<
-0.Q1
..0,02
50
I
20
,.,,,
<:
;<""W'.
15
10
5
Time (103 Years)
'-
,
0
0
Maximum
"""':<;",TI
6,
x
<1>
o
Q)
Q.:
0.02 ,
200
,...----...."
~~i£
c
Q
r
0.01
150 ~
tf'
'"
~;
0)
200
<1>
o
~
Q.: -0.01
2-
-G.O?
I
20
./
'--./
'-"
15
10
5
Time (103 Years)
I
0
100
s;-
50
(l)
0
<
2-
Supplementary information to "Human Evolution and Migration in a Variable
Environment: The Amplifier Lakes of East Africa" by Martin H. Trauth, Mark A.
Maslin, Andreas G.N. Bergner, Alan Deino, Annett Junginger, Eric Odada, Daniel O.
Olago, Lydia Olaka, Manfred R. Strecker
In the following, we provide a more detailed compilation of the volcano tectonic and fluvio-lacustrine
history of major lake basins from the North to South (Suppl. Figs I and 2).
Volcano-tectonic evolution of the East African rift basins - The earliest evidence for the development of
the East African Rift System (EARS) is an intense magmatic activity in Ethiopia between 45 and 35 Ma and
in northern Kenya at around 33 Ma that continued to about 25 Ma, whereas the magmatic activity of the
central and southern segments of the rift in Kenya and Tanzania started between 15 and 8 Ma (e.g., Ebinger
et aI., 1993). The driver for the magmatic activity as well as for the regional plateau uplift is the intrusion of
an asthenopheric plume in East Africa (Ebinger et al., 1993). The exact timing of the development of the East
African and Ethiopian plateaus is intensely discussed and ages for earliest topographic uplift range from 20
Ma (Pik et al., 2006; 2008) to 8 Ma (Spiegel et al., 2007). The tectonic uplift, however, was the first evidence
for an climatic significance of the EARS and inspired the discussion about an influence of the plateau
Formation, regional aridification and vegetation changes (e.g., Dart. J953; Potts, 1998; Sepulchre et al..
2006). The plateau uplift was associated with an enormous magmatic activity in the area. In the Ethiopian
Rift, volcanism started between 45 and 33 Ma, in northern Kenya at around 33 Ma and continued to about 25
Ma, whereas the magmatic activity of the central and southern segments of the rift in Kenya and Tanzania
started between IS and 8 Ma (Bagdasaryan ct al.. 1973; Crossley. 1979; Davidson and Rex. 1980; Crossley
and Knight, 198J; McDougall and Watkins, 1987; Morley et aI., 1992: Dawson. 1992: Ebinger et al.. 1993:
George et al., 1998).
While these early stages of rifting were characterized by updoming and downwarping in the later rift sectors,
faulting progressed from north to south. Major faulting in Ethiopia between 20 to 14 Ma was followed by the
generation of east-dipping faults in Kenya between 12 and 7 Ma on NNW striking and E dipping normal
faults during SW-SE extension, creating a half graben with a rollover monocline (Williams et al., 1983;
Baker et a!., 1988, Blisniuk and Strecker, 1990; Strecker and Bosworth, 1991; Ebinger et al., 2000). These
early half grabens were subsequently faulted antithetically between about 5.5 and 3.7 Ma during a slightly
rotated stress field with WSW-ESE orientation, which generated a full-graben morphology (Baker et al.,
1988, Blisniuk and Strecker. 1990). The earliest evidence for lakes in a half/early-full graben setting is a -90
m thick diatomite sequence that overlies -4.6 Ma old trachytic lava flows and is overlain by 4.5-3.4 Ma old
Kinangop tuffs exposed in the valley of the Turasha River on the present Kinangop Plateau East of the
modem Naivasha basin (Trauth et al., 2007). The diatom assemblages indicate a shallow freshwater
environment rather than the conditions in a large and deep lake (Trauth et al., 2007). In cannot excluded,
however, that the Turasha deposits represent the sediments of larger, but relatively shallow lake that
expanded towards the North and South along the axis of the rift.
Prior to the full-graben stage, the large Aberdare volcanic complex with elevation in excess of 4,000 m was
established (Baker et al.. 1972; Williams et al.. 1983) (Suppl. Fig. I). Since then, the Aberdare range in the
East of the modern Naivasha basin together with the Mau Escarpment, more than 3,000 m high, to the West
form important orographic barriers in Kenya that form a rainshadow for moisture-bearing winds from both
the Indian Ocean and the Atlantic Ocean via the Kongo basin. Between 3.6 and 2.7 Ma, ongoing WSW-ESE
extension created W- to WSW-dipping normal faults with dextral components creating the Aberdare
escarpment along the Sattima fault (Strecker et al., 199 J). By 2.6 Ma, the graben was further segmented in
the Central Kenya Rift by West dipping faults, creating the 30 km wide intrarift Kinangop Plateau and the
tectonically active 40-km-wide inner rift (Baker et al., 1988; Strecker ct aI., 1990). The central graben
depression in the Naivasha area between the Kinangop Plateau and the Mau escarpment went through a
phase of subsidence after 2.6 Ma before voluminous trachyte eruptions occurred that did not overlap with the
plateau after 1.9 Ma during W-E extension. The inner rift was subsequently covered by trachytic, basaltic,
and rhyolithic lavas and continues to be affected by normal faulting leading to further segmentation of the
rift floor (Strecker et aI., J 990; Bosworth and Strecker, 1997). There is no unambiguous evidence from
sedimentation patterns in the Magadi-Narron and Olduvai basins, however, that major fault-bounded basins
existed at that time (Foster ct al., 1997). Major normal faulting in these regions occurred at 1.2 Ma and
produced the present-day rift escarpments (Foster et aI., 1997). Subsequent Late Quaternary structural enechelon segmentation during west-northwest to east-southeast oriented extension created numerous subbasins in the individual rift sectors that repeatedly hosted smaller lakes (Baker ct al., 1988).
The magmatic and tectonic evolution of the Kenyan rift segments shows that conditions for the
establishment of large lakes existed by approximately 3 Ma. However, continued faulting and en-echelon
segmentation during WNW-ESE oriented extension created numerous smaller sub-basins that repeatedly
hosted lakes during the Pleistocene (Strecker et al., 1990; Bosworth and Strecker, 1997).
Examples for variable environments on all time scales - East African lakes fluctuated on all time scales
(SUppI. Fig. 2). The chronology of Plio-Pleistocene lake-level variations in East Africa indicates consistent
wetter and more seasonal conditions every ca. 0.8 Ma, with periods of precessional-forced extreme climate
variability at ca. 4.7-4.3 Ma, 4.2-3.8 Ma, -3.6-3.4 Ma, -3.2-2.90 Ma, 2.7-2.5 Ma, 1.9-1.7 Ma and 1.1-0.9 Ma
and after ca. 0.2 Ma before present (Trauth et al., 2003, 2005, 2007; Maslin and Trauth, 2(09). On shorter
time scales, the history of these large lakes reveal rapid (less than one thousand years) and large-magnitude
fluctuations (up to -300 meters, Garcin et al., 2009). The internal variability of the large-lake episodes is best
documented for the Plio-Pleistocene Chemeron Formation (3.5 to 1.7 Ma) in the Tugen Hills (Deino et al.,
2006; Kingston et al., 2007), the Mid Pleistocene Olorgesailie Formation in the Southern Kenya Rift (1.2 to
0.5 Ma) (Behrensmeyer et al., 2002; Owen et aI., 2008), and the Late Pleistocene 01 Njorowa Gorge
Formation in the Central Kenya Rift «0.2 Ma) (Trauth et aI., 2001, 20(3). The revised analyses of terrestrial
dust records from marine sediments by Trauth et aI. (2009) has recently shown that African climate is mostly
influenced by low-latitude climate transitions such as the closure of the Indonesian Seaway (Cane and
Molnar, 200 I) and associated reduced heat flow to the tropics and the onset of the Walker Circulation near
1.9 Ma (Ravelo et al., 2005; Trauth et al., 2009) rather than the intensification of the Northern Hemisphere
glaciations as a result of the closure of the Panama Isthmus (deMenocal, 1995, 2004).
Recent work by Andrew Hill, John Kingston and colleagues on the upper Chemeron Formation (3.5 to 1.7
Ma) exposed in the Tugen Hills (western Baringo-Bogoria Basin) revealed a sequence of five major
diatomites (Dei no et aI., 2006; Kingston et al., 2007) (Suppl. Fig. 2a). Single-crystal
40 Arj39Ar
determinations show they occur between 2.66 and 2.56 Ma at -20 kyr intervals (Dcino ct al., 2006). Detailed
mapping and stratigraphic analysis suggest that these cycles with precessional periodicity are not due to local
tectonic movements but reflect climate changes (Dei no et al., 2006). First investigations of the diatom flora
contained in these diatomites indicate the existence of large deep lakes between 2.66 and 2.56 Ma (Owen,
2002). The lake episode recorded in the Chemeron Formation correlates with several important lake episodes
observed in the Afar Basin and the Ethiopian Rift (Williams et al., 1979).
The Olorgesailie Formation has been intensely studied by the team of Rick Potts since the 80's (e.g., Potts,
1999; Behrensmeyer et aI., 2002; Owen et al., 2008) (Suppl, Fig. Zb). The Olorgesailie Formation records the
Formation of a closed-basin environment shortly before 0.992 Ma, alternating between lacustrine and
subaerial conditions for a period of -500 kyr but no episodes of major erosion (Behrensmeyer et al., 2002;
Owen et al., 2008). The most important lake period occurred between 0.992 and 0.974 Ma as documented by
the deposition of the main diatomite bed (Behrensmeyer et aI., 2002; Owen et aI., 2008). After 0.500 Ma, the
basin experienced phases of valley cutting associated with base-level fluctuations of around 50 mover
periods of 104 to 105 years (Potts, 1988; 1998; Behrensmcyer et al., 2002; Owen ct al., 2008). The
stratigraphy of the Olorgesailie Formation, however, has recently been the matter of discussions (see Trauth
et al., 2005; Owen et al., 2008; Trauth and Maslin, 2009; Owen et aI., 2009). According to the newest
interpretation by Owen et al. (2008), the Olorgesailie basins has no environmental record during much of the
time between 1.2 and 0.49 Ma. Owen et al. (2008) proposed relatively short periods of full lacustrine
conditions, whereas soil formation, reworking and erosion are the predominant processes during much of the
time between 1.2 and 0.49 Ma. Instead, we propose a more straightforward interpretation of the Olorgesailie
record based on the raw stratigraphic evidence and lake-level record as published by Behrensmeyer et al.
(2002), the Ar/Ar chronology published by Deino et al. (1990; 1992) and the additional Ar/ Ar age of 1.2 Ma
near the base of the section provided by Owen ct al. (2008) (Suppl. Fig. 2b). A robust stratigraphy
independent from any assumptions about sedimentation rates, the character of the deposits and the
occurrence of reworked or eroded material was obtained by simple linear interpolation of the lake-level
record Behrensmeyer et al. (2002) taking into account the Ar/Ar ages by Deino et ill. (1990; 1992) and Owen
et al. (2008). The large error bar of the Ar/Ar age near 1.0 Ma and the large undated interval between -1.2
and 1.0 Ma introduces some degree of uncertainty that allows to shift the earliest lake episode by -0.1 Ma
towards older times. The lake-level record in the Olorgesailie basin then perfectly matches the earth's
eccentricity cycle and therefore suggest that indeed a large, but fluctuating lake occurred in the southern
Kenya rift during a relatively long period of time. As already stated in Trauth et al. (2005, 2007), however,
the lake-level was subjected to considerable fluctuations all the time, as also indicated by the analysis of
diatom assemblages (Owen ct al., 2008). The lake episode recorded in the Olorgesailie Formation also
correlates nicely with fluctuating paleo-lakes in the Central Kenya Rift (Trauth et (II., 2005, 2(07), the
Suguta Valley (Dunkley et al., 1993), the Omo-Turkana basin (Feibel et al., 1991) and in the Afar Basin
(Gasse,1990).
The 01 Njorowa Gorge Formation in the southern Naivasha basin shows five highstands and intermittent
lowstand recurring at half-precessional (ca. II kyr) cyclicities (Trauth et a1., 2001. 2003) consistent with the
concept of low-latitude orbital forcing and climate (Berger et al., 1997, 2(06) (Suppl.T'ig. 2c). The Late
Pleistocene Lake Naivasha was characterized by the deposition of up to 4 m thick diatomites containing a
mixed planktonic and benthic-epiphytic diatom community that indicate alternating freshwater and highly
alkaline conditions (Trauth et al., 200 1, 2003). Additional evidence for the paleo-water depths of these young
lakes comes from the altitude of paleo-shorelines and sediment outcrops (Trauth et al., 2001, 2003). Basin
reconstructions and hydrological modeling suggests that the highest lake levels reached ca. 150 m during the
peak of the wet episode between ca. 145 and 60 kyr before present (Bergner et al.. 2003, 2004). The 01
Njorowa Gorge Formation also contains evidence for remarkable droughts in the Central Kenya Rift similar
to the ones reported from the Malawi basin by Brown et a!. (2007). The corresponding sediments contain
authigenic silicates such as zeolites that document chemical weathering of silicic volcanic glass in an
extremely alkaline lake environment. No lake, however, indicates the complete absence of lake sediments.
The sedimentary record of basin contains fluvial or eolian deposits, and primarily deposited, impact craters
of pumice lapilli on mud surfaces or reworked pyroclastic material. The lake episode recorded in the 01
Njorowa Gorge Formation also correlates with the Kibish Formation in the Omo along the Omo River
(Butzer et al., 1969; Brown et al., 2(08). Uranium-series ages for hydrothermal deposits recording
geothermal activity in the Suguta Valley (Northern Kenya Rift) suggest elevated water tables and increased
availability of meteoric water associated with more humid climate as early as 133±11 kyr BP (Sturchio et aI.,
1993). High lake levels can be inferred from shorelines in the Magadi-Natron Basin in the southern part of
the rift, where littoral stromatolites provide U-series ages of 135 kyr BP (Hillaire-Marcel et a!., 1986).
The tenninal Pleistocene/Early
Holocene sediments and shorelines of Lakes Naivasha and Nakuru
Elmenteita are also well-studied lakes for the last precessional cycle and associated high-lakes levels
(Washbourn-Kamau, 1970; 1975; 1977; Richardson and Dussinger.1986: Trauth et al., 2003; Duhnforth et
al., 20(6) (Suppl. Fig. 2d). Between ca. 13 and 5 kyr before present the lake level was I J 0 m (Lake
Naivasha) and 180 m (Lake Nakuru-Elmenteita) above the modern lakes. High lakes with similar amplitudes
were also reported from many other East African lake basins, such as Lake Abhe (+ J 60 m) (Gasse et al.,
1977), Lake Ziway (+112 m) (Gillespie et al., 1983), Lake Suguta (+280 m) (Garcin et al., 2009), Lake
Turkana (+80 m) (Owen et al., 1982; Hastenrath and Kutzbach, 1983: Vincens et al., J 989), Lake NakuruElmenteita (+180 m) (Washbourn-Kamau,
J 967; 1970), Lake Naivasha
(+ 100 m) (Wash bourn, 1975;
Hastenrath and Kutzbach, 1983) and Lake Rukwa (+200 m) (Kervyn et al., 20(6) that mark the so-called
African Humid Period (spanning ca. 15,500-5,500 cal. yrs BP) (c.g., Gassc, 2000; Barker et al., 2004). The
abruptness of the onset and termination of the African Humid Period and associated environmental shifts is a
matter of debate (deMenocal et al., 2000; Kropelin et al., 2008a, 2008b; Brovkin and Claussen, 2008;
deMenocal, 2008; Cole et al., 2(09). Also intensely debated is the relative importance of high-latitude glacial
shifs and reduced thermohaline circulation during Heinrich and Younger Dryas events during that time
interval (Trauth et a!., 2003; Brown et al., 2007; Tierney et al., 2008; Garcin et al., 2009).
During the last millenium, Lake Naivasha fluctuated considerably by ~30 m during decades (Verschuren et
al., 2000) (Suppl. Fig. 2e). The water level variations were reconstructed using sediment stratigraphy, diatom
assemblages and fossil midge assemblages. According to these reconstructions, East Africa as significantly
drier than today during the Medieval Warm Period (MWP, AD 1000-1270) and wetter during the Little Ice
Age (LlA, AD 1270-1850) which was interrupted by three prolonged dry episodes. The humidity changes in
the Naivasha basin correlate with long-term changes in solar radiation suggesting that decade-scale rainfall
variations are largely controlled by solar forcing (Verschuren et al., 2000). The Naivasha record as a record
of climate changes during the last millenium, however, was also subject to intense discussions (see summary
in Gasse et al., 2002). Six ice cores from Kilimanjaro providing an ~I I .7-thousand-year record of Holocene
climate and environmental variability for eastern equatorial Africa suggest substantial cooling (Thompson et
al., 2002). These and other records of East African climate changes during the last millennium illustrate the
spatial complexity of environmental changes on relatively short times scales (e.g., Gasse et al., 2002;
Thompson ct al., 2006; Russell et al., 2007).
The modern lake-level fluctuations of Lake Naivasha based
range ofa few (1 to 6) meters (Data from Ministry of Water
1979; Ase, 1987). The present climate patterns in East
convergence zones, superimposed upon regional factors
on satellite altimetry and observations are in the
and Irrigation of Kenya. 2008, and Vincent et al.,
Africa include several major air streams and
associated with topography, large lakes, and
maritime influence (Nicholson, 1996). Regional wind and pressure patterns include the Congo air stream
with westerly and southwesterly air flow, as well as the NE and SE monsoons. In contrast to the Asian SW
monsoon, both monsoons over East Africa are relatively dry. In contrast, the Congo air stream is humid and
associated with rainfall. These major air streams are separated by the Intertropical Convergence Zone (ITCZ)
and the Congo Air Boundary. Rainfall in East Africa is mainly linked to the passage of the ITCZ causing a
strongly bimodal annual cycle (Nicholson, 1996). The period during April-May is considered to be the 'long
rains' while the October-November period is called the 'short rains', following the latitudinal position of the
overhead sun with a time lag of about 4-6 weeks. Fluctuations in the intensity of precipitation are linked to
E-W adjustments in the zonal Walker circulation associated with the Indian Ocean Dipole and the EI Nino/
Southern Oscillation (Saji et aI., 1999; Moy et aI., 2002) (Suppl. Fig. 21).
References and Notes
Ase, L.E., 1987. Water level variations of Lake Naivasha. Geografiska Annaler 69A, 415-429.
Bagdasaryan, G.P., Gerasimovskiy, Vl., Polyakov, A.I., Gukasyan, R.K., 1973. Age of volcanic rocks in the
rift zones of East Africa. Geochemistry International 10, 66-71.
Baker, B.H., Mohr, P.A., Williams, L.A.J., 1972. Geology of the eastern rift system of Africa.
Baker, B.H., Mitchell, J.G., Williams, L.A.J., 1988. Stratigraphy, geochronology and volcano-tectonic
evolution of the Kedong-Naivasha-Kinangop
region, Gregory Rift Valley, Kenya. Journal of the
Geological Society 145, 107-116.
Barker, P.A., Talbot, M.R., Street-Perrott, FA., Marret, F, Scourse, J.D., Odada, E., 2004. Late Quaternary
climatic variability in intertropical Africa. In: Battarbee, R.W., Gasse, F, Stickley, C.E. (Ed.), Past
Climate Variability through Europe and Africa. Developments in Paleoenvironmental Research. Kluwer
Academic Publishers, Dordrecht, pp. 117-138.
Behrensmeyer, A.K., Potts, R., Deino, A., Ditchfield, P., 2002. Olorgesailie, Kenya: a million years in the life
ofa rift basin. SEPM Special Publication 73, 97-106.
Berger, A., Loutre, M.F, 1991. Insolation values for the climate of the last 10 million of years. Quaternary
Sciences Review 10,297-317.
Bergner, A., Trauth, M., 2004. Comparison of the hydrological and hydrochemical evolution of Lake
Naivasha (Kenya) during three highstands
between 175 and 60 kyr BP Palaeogeography, Palaeoclimatology, Palaeoecology 125, 17-36.
Bergner, A.G.N., Trauth, M.H., Bookhagen, B., 2003. Magnitude of precipitation / evaporation changes in
the Naivasha Basin (Kenya) during the last 150 kyrs. Global and Planetary Change 36, 117-136.
Bergner, A.G.N., Strecker, M.R., Trauth, M.H., Deino, A., Gasse, F, Blisniuk, P., Duhnforth, M, in press.
Tectonic versus climate influences on the evolution of the lakes in the Central Kenya Rift. Quaternary
Science Reviews.
Bosworth, W., Strecker, M.R., 1997. Stress field changes in the Afro-Arabian rift system during the Miocene
to Recent period. Tectonophysics 278, 47-62.
Boven, A., 1992. The applicability of the K-Ar and 40Ar/39Ar techniques to Quaternary volcanics: a
fundamental and experimental investigation. Vrije Universiteit Brussel, Brussel.
Brovkin, v., Claussen, M., 2008. Comment on "Climate-Driven Ecosystem Succession in the Sahara: The
Past 6000 Years". Science 322, 1326b-c.
Brown, E.T., Johnson, T.c., Scholz, C.A., Cohen, A. S. King, J. W., 2007. Abrupt change in tropical African
climate linked to the bipolar seesaw over the past 55,000 years. Geophysical Research Letters 34,
L20702.
Brown, FH., Feibel, C.S., 1991. Stratigraphy, depositional environments, and palaeogeographiy of the Koobi
Fora Formation. In: Harris, J.M. (Ed.), Koobi Fora Research Project. Clarendon, Oxford, pp. 1-30.
Brown, FH., Fuller, C.R., 2008. Stratigraphy and tephra of the Kibish Formation, southwestern Ethiopia.
Journal of Human Evolution 55, 366-403.
Butzer, K.W., Brown, FW., Thruber, D.L., 1969. Horizontal sediments of the lower Omo valley; the Kibish
formation. Quaternaria II, 15-29.
Cane, M.A., Molnar, P., 2001. Closing of the Indonesian seaway as a precursor to east African aridification
around 3-4 million years ago Nature 411, 157-162.
Casanova, J., 1986. Les stromatolites continentaux: paleoecologie, paleohydrologie, paleoclimatologie.
Application au Rift Gregory. Universite d'Aix-Marseille II, Marseille.
Clarke, M.C.G., Woodhall, D.G., Allen, D., Darling, G., 1990. Geological,
volcanological
and
hydrogeological controls on the occurence of geothermal activity in the area surrounding Lake Naivasha,
Kenya. BGS, Nairobi.
Crossley, R., 1979. The Cenozoic stratigraphy and structure of the western part of the Rift Valley in southern
Kenya. Journal of the Geological Society London 136,393-405.
Crossley, R., Knight, R.M., 1981. Volcanism in the western part of the rift valley in southern Kenya. Bulletin
of Volcanology 44, 117-128.
Davidson, A., Rex, D., 1980. Age of volcanism and rifting in southwestern Ethiopia. Nature 283, 657-659.
Dawson, J.B., 1992. Neogene tectonics and volcanicity in the North Tanzania sector of the Gregory Rift
Valley: contrasts .... Tectonophysics 204, 81-92.
Deino, A., Potts, R., 1990. Single-crystal 40 Ar139 Ar dating of the Olorgesailie Formation, Southern Kenya
Rift. Journal of Geophysical Research 95, 8453-8470.
Deino, A.L., Kingston, J.D., Glen, J.M., Edgar, R.K., Hill, A., 2006. Precessional forcing of lacustrine
sedimentation in the late Cenozoic Cherneron Basin, Central Kenya Rift, and calibration of the Gauss/
Matuyama boundary. Earth and Planetary Science Letters 247, 41-60.
deMenocal, P., 1995. Plio-Pleistocene African Climate. Science 270,53-59.
deMenocal, P., 2000. Abrupt onset and termination of the African Humid Period: rapid climate responses to
gradual .... Quaternary Science Reviews 19,347-361.
deMenocal, P., 2004. African climate change and faunal evolution during the Pliocene-Pleistocene. Earth and
Planetary Science Letters 220, 3-24.
Duhnforth, M., Bergner, A.G.N., Trauth, M.H., 2006. The hydrological budget of the Nakuru-Elmenteita
basin, Central Kenya Rift, during the Early Holocene wet period. Journal of Paleolimnology 36,281-294.
Dunkley, P.N., Smith, M., Allen, D.J. and Darling, w.G., 1993. The geothermal activity and geology of the
northern sector of the Kenya Rift Valley. British Geological Survey Research Report. British Geological
Survey, Nottingham, p. 185.
Ebinger, CJ., Rex, D.C., Yemane, T, Kelley, S., 1993. Volcanism and extension between the Main Ethiopian
and Gregory rifts. In: Thorweihe, U., Schandelmeier, H. (Ed.), Geoscientific Research in Northeast
Africa, Rotterdam, pp. 301-304.
Ebinger, CJ., Sleep, N.H., 1998. Cenozoic magmatism throughout east Africa resulting from impact of a
single plume. Nature 385, 788-791.
Ebinger, c.r., Yemane, T, Harding, DJ., Tesfaye, S., Kelley, S., Rex, D.C., 2000. Rift deflection, migration,
and propagation: Linkage of the Ethiopian and Eastern rifts, Africa. Bulletin of the Geological Society of
America 112, 163-176.
Evemden, lG., Curtis, G.H., 1965. The potassium-argon dating of late Cenozoic rocks in East Africa and
Italy. Current Anthropology 6,177-189.
Foster, A., Ebinger, c., Mbede, E., Rex, D., 1997. Tectonic development of the northern Tanzanian sector of
the east African rift system. Journal of the Geological Society London 154, 689-700.
Garcin, Y, Junginger, A., Melnick, D., Olago, D.O., Strecker, M.R., Trauth, M.H., 2009. Late PleistoceneHolocene rise and collapse of Lake Suguta, northern Kenya Rift. Quaternary Science Reviews 28,
911-925.
Gasse, 2002. Kilimanjaro's Secrets Revealed. Science 298, 548-549.
Gasse, F., 1977. Evolution of Lake Abhe (Ethiopia and TFAI), from 70,000 b.p. Nature 265, 45-47.
Gasse, F., 1990. Tectonic and climatic controls on lake distribution and environments in Afar from Miocene
to present. In: Katz, BJ. (Ed.), Lacustrine Basin Exploration, Case Studies, and Modern Analogs. AAPG
Memoir, pp. 1-34.
George, R.M.M., Rogers, N.W., Kelley, S., 1998. Earliest magmatism in Ethiopia: evidence for two mantle
plumes in one flood basalt province. Geology 26, 923-926.
Gillespie, R., Street-Perrott, F.A., Switsur, R., 1983. Post-glacial arid episodes in Ethiopia have implications
for climate prediction. Nature 306, 680-683.
Hastenrath, S., Kutzbach, lE., 1983. Paleoclimatic Estimates from Water and Energy Budgets of East
African Lakes. Quaternary Research 19, 131-153.
Hay, R.L., 1976. Geology of the Olduvai Gorge. A study of sedimentation in a semiarid Basin. University of
California Press, Berkeley, Los Angeles, London.
Hay, R.L.. Kyser, TK., 2001. Chemical sedimentology and paleoenvironmental history of Lake Olduvai, a
Pliocene lake in northern .... Bulletin of the Geological Society of America 113, 1505-1521.
Hillaire-Marcel, c., Carro, 0., Casanova, L, 1986. 14C and Th/u Dating of Pleistocene and Holocene
Stromatolites from East African Paleo lakes Quaternary Research 25, 312-329.
Kervyn, F., Ayub, S., Kajara, R., Kanza, E., Temu, B., 2006. Evidence of recent faulting in the Rukwa rift
(West Tanzania) based on radar interferometric DEMs. Joumal of African Earth Sciences 44, 151-168.
Kingston, J., Deino, A., Hill, A., 2007. Astronomically forced climate change in the Kenyan Rift Valley 2.72.55 Ma: implications for the evolution of early hominin ecosystems. Joumal of Human Evolution 53,
487-503.
Kropelin, S., Verschuren, D., Lezine, A.-M., 2008. Response to Comment by Brovkin and Claussen on
"Climate-Driven Ecosystem Succession in the Sahara: The Past 6000 Years". Science 322, 1326.
Kropelin S., V.D., Lezine A.-M., Eggermont H., Cocquyt c., Francus P., Cazet 1.-P., Fagot M., Rumes B.,
Russell 1. M., Darius F., Conley D. 1., Schuster M., Suchodoletz H. V., Engstrom D. R., 2008. ClimateDriven Ecosystem Succession in the Sahara: The Past 6000 Years. Science 329, 765-768.
Leakey, M.G., Feibel, C.S., McDougall, 1., Walker, A., 1995. New four-million-year-old hominid species
from Kanapoi and Allia Bay, Kenya. Nature 376, 565-571.
Leakey, M.G., Feibel, C.S., McDougall, 1., Ward, c., Walker, A., 1998. New specimens and confirmation of
an early age for Australopithecus anamensis. Nature 393, 62-66.
Leblanc, M., Favreau, G., Maley, 1., Nazoumou, Y., Leduc, c., Stagnitti, F., van Oevelen, P., Delclaux, F.,
Lemoalle, J., 2006. Reconstruction of Megalake Chad using Shuttle Radar Topographic Mission data.
Palaeogeography, Palaeoclimatology, Palaeoecology 239, 16-27.
Lepre, c.J., Quinn, R.L., Jcordens, lC.A., Swisher III, c.c., Feibel, C.S., 2007. Climatic forcing of
terrestrial environmental change during the emergence of H. erectus: evidence from the KBS Member of
the Koobi Fora Formation. Journal of Human Evolution 53, 504-514.
Macdonald, R., Rogers, N.W., Fitton, 1.G., Black, S., Smith, M., 2001. Plume-Lithosphere Interactions in the
Generation of the Basalts of the Kenya Rift, East Africa. Journal of Petrology 42,877-900.
McCall, GJ.H., Baker, B.H., Walsh, L, 1967. Late Tertiary and Quaternary sediments of the Kenya Rift
Valley. In: Bishop, W.w., Clark, lD. (Ed.), Background to evolution in Africa. University of Chicago
Press,pp.191-220.
McDougall, I., 1985. K-Ar and 40Ar/39Ar dating of the hominid-bearing Pliocene-Pleistocene sequence at
Koobi Fora, Lake Turkana, northern Kenya. Geological Society of America Bulletin 96, 159-175.
McDougall, I., Watkins, R.T., 1988. Potassium-Argon ages of volcanic rocks from northeast of Lake
Turkana, northern Kenya. Geological Magazine 125, 15-23.
Morley, CX., Wescott, W.A., Stone, D.M., Harper, R.M., Wigger, S.T., Karanja, F.M., 1992. Tectonic
evolution of the northern Kenya rift. Journal of the Geological Society London 149,333-348.
Morley, CX., 1994. Interaction of deep and shallow precesses in the evolution of the Kenya rift.
Tectonophysics 236, 81-91.
Nicholson, S.E., 1996. A review of climate dynamics and climate variability in Eastern Africa. In: TC.
Johnson, E.O.O. (Ed.), The Limnology, Climatology and Paleoclimatology of the East African Lakes, The
International Decade for the East African Lakes (IDEAL). Gordon and Breach, Newark, NJ, pp. 25-56.
Owen, 2002. Sedimentological Characteristics and Origins of Diatomaceous Deposits in the East African
Rift System. SEPM Special Publication 73, 233-246.
Owen, R.B., Potts, R., Behrensmeyer, AX., Ditchfield, P., 2008. Diatomaceous sediments and environmental
change in the Pleistocene Olorgesailie Formation, southern Kenya Rift Valley. Palaeogeography,
Palaeoclimatology, Palaeoecology 269, 17-37.
Party, K.W., 1991. Large scale variations in lithospheric structure along and across the Kenya Rift. Nature
317,130-134.
Pik, R., Marty, B. Hilton, D.R., 2006. How many mantle plumes in Africa? Geochemical point of view.
Chemical Geology 226, 100-114.
Pik, R., Marty, B., Carignan, 1., Yirgu, G. and Ayalew, T, 2008. Timing of East African Rift development in
southern Ethiopia: Implication for mantle plume activity and evolution of topography. Geology 36,
167-170.
Potts, R., 1998. Environmental hypotheses of hominin evolution. Yearbook of Physical Anthropology 41,
93-136.
Potts, R., 1999. Paleolandscape variation and Early leistocene hominid activities: Members I and 7,
Olorgesailie Formation, Kenya. Journal of Human Evolution 37, 747-788.
Prodehl, c., Ritter, J.R.R., Mechie, J., Keller, G.R., Khan, M.A., Jacob, B., Fuchs, K., Nyambok, 1.0., Obel,
J.D., Riaroh, D., 1997. The KRISP 94 lithospheric investigation of southern Kenya-the
experirnents and
their main results. Tectonophysics 278,121-147.
Ravelo, c., Andreasen, D., Olivarez, A., Lyle, M., Wara, M., 2004. Regional climate shifts caused by gradual
global cooling in the Pliocene epoch. Nature 429,263-267.
Richardson, lL., Dussinger, R.A, 1986. Paleolimnology of mid-elevation lakes in the Kenya Rift Valley.
Hydrobiologia 143,167-174.
Russell, J., Verschuren, D., Eggerrnont, H., 2007. Spatial complexity of 'Little Ice Age' climate in East
Africa: sedimentary records from two crater lake basins in western Uganda. The Holocene 17, 183-193.
Saji, N.H., Goswami, B.N., Vinayachandran, P.N., Yamagata, T, 1999. A dipole mode in the tropical Indian
Ocean. Nature 401,360-363.
Schreck, c., Semazzi, F., 2004. Variability of the recent climate of eastern Africa. International Journal of
Climatology 24, 681-701.
Sepulchre, P., Ramstein, G., Fluteau, F., Schuster M., Tiercelin, U., Brunet, M., 2006. Tectonic Uplift and
Eastern Africa Aridification. Science 313, 1419- I423.