Aeolian Research 8 (2013) 65–73 Contents lists available at SciVerse ScienceDirect Aeolian Research journal homepage: www.elsevier.com/locate/aeolia Spatial patterns of aeolian sediment deposition in vegetation canopies: Observations from wind tunnel experiments using colored sand Katrin Suter-Burri a,b,⇑, Christof Gromke a,c, Katherine C. Leonard a,d, Frank Graf a a WSL Institute for Snow and Avalanche Research SLF, CH-7260 Davos Dorf, Switzerland Institute for Integrative Biology IBZ, ETH Zurich, CH-8092 Zurich, Switzerland c Building Physics and Services, Eindhoven University of Technology, 5600 MB Eindhoven, The Netherlands d Cooperative Institute for Research in Environmental Sciences, University of Colorado at Boulder, 216 UCB, Boulder, CO 80309, USA b a r t i c l e i n f o Article history: Received 28 July 2012 Revised 4 November 2012 Accepted 15 November 2012 Available online 12 December 2012 Keywords: Wind tunnel Vegetation Colored sand Erosion a b s t r a c t In environments affected by wind erosion, plants act as traps for aeolian sediment, which leads to a small-scale mosaic of depositional and erosional sediment transport regimes. This wind tunnel study used colored sand to visualize spatial patterns of sediment redistribution within grass canopies. Wind tunnel experiments were performed with high-, medium- and low-density canopies of Lolium perenne, corresponding to vegetation covers of 47%, 16% and 4%, respectively. In the low and medium-density canopies, the wake areas downstream of the tussocks were the primary locations of sediment deposition. In the medium-density canopy, these wedge-shaped wake deposits overlapped with the adjacent downstream tussocks, while in the low-density canopy they did not, indicating that these vegetation densities respectively represented wake-interference and isolated roughness flow. In the high-density canopy, very few sand grains were entrained by the wind, and were mostly deposited within the disturbed zones surrounding the tussocks. The deposited grains were evenly distributed around the tussocks in the highdensity canopy without pronounced accumulations on their upstream, downstream or lateral sides. We interpret the high-density canopy as a skimming flow aerodynamic regime. The fraction of the sand surface which was exposed to erosion was substantially smaller than the area not covered by grasses. It accounted for 67–78% of the non-covered surface in the low-density canopy, and for 44–77% of the surface in the medium-density canopy. This finding indicates that wind erosion models overestimate the sediment source area if they assume the erodible area is the entire exposed surface not covered by roughness elements. Ó 2012 Elsevier B.V. All rights reserved. 1. Introduction Vegetation affects soil erosion by the wind at multiple spatial and temporal scales. The above-ground parts of vegetation act as non-erodible roughness elements that modify the near-surface wind, while their roots help stabilize the soil. The effect of vegetation on aeolian sediment transport is attributed to three mechanisms (Wolfe and Nickling, 1993): (i) surface sheltering, (ii) momentum extraction from the wind, and (iii) trapping of windborne sediment. Surface sheltering is provided both by covering a portion of the ground surface and limiting the area of bare ground available to be eroded and by the wakes of reduced mean wind velocity downflow from a plant. Morris (1955) defined three flow regimes depending on the wake development and the proportion of protected to total ⇑ Corresponding author at: WSL Institute for Snow and Avalanche Research SLF, CH-7260 Davos Dorf, Switzerland. Tel.: +41795747614. E-mail addresses: [email protected] (K. Suter-Burri), [email protected] (C. Gromke), [email protected] (K.C. Leonard), [email protected] (F. Graf). 1875-9637/$ - see front matter Ó 2012 Elsevier B.V. All rights reserved. http://dx.doi.org/10.1016/j.aeolia.2012.11.002 surface area: (i) isolated roughness flow, where there is no interaction between wakes and adjacent downstream roughness elements, (ii) wake interference flow, where wakes from upstream elements intercept downstream elements, and (iii) skimming flow, where wakes completely overlap and the entire ground surface is sheltered (Fig. 1). The flow regime depends on both the size and spacing of roughness elements and on the free-stream or mean wind speed, and is a critical factor in determining the amount of wind erosion a surface will experience (Morris, 1955; Lee and Soliman, 1977; Wolfe and Nickling, 1993; Breshears et al., 2009). Momentum extraction from the wind by vegetation reduces the erosive force of the wind. By absorbing a part of the total shear stress of the wind, plants decrease the shear stress acting on the ground surface. Trapping of sediment by vegetation occurs through two mechanisms. Windborne particles can be removed from the air stream when they hit plant surfaces, or when the wind velocity becomes too low in the immediate vicinity of plants. By these means, plants cause particles to settle from the flow, resulting in characteristic 66 K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 Fig. 1. Schematic representation of the three flow regimes defined by Morris (1955). Figure modified after Oke (1988) and Gromke (2008). deposition patterns within vegetation canopies. A landscape experiencing wind erosion is thus a small-scale mosaic of depositional and erosional areas (Okin et al., 2006). These sediment redistribution processes have important implications for the development of a landscape’s surface morphology and for soil resource availability. Wind erosion also depletes nutrients from plant interspaces and accumulates them within and around plants, a process known as the resource island phenomenon (Bielders et al., 2002; Li et al., 2007). Several methods have been used to study small-scale spatial patterns of aeolian sediment deposition. Udo and Takewaka (2007) used a laser displacement sensor to measure wind-induced bed elevation changes in flexible artificial plant canopies in a wind tunnel setting. They found that canopies of low height, high density and high flexibility promoted bed accumulation and reduced sand transport downstream of the canopy. In several field studies, small-scale erosion and deposition processes have been monitored by marking the changing level of the soil surface on steel rods driven into the ground (Wiggs et al., 1995; Hesse and Simpson, 2006; Li et al., 2008). Other field studies have measured windborne sediment mass fluxes in heterogeneous plant canopies, indirectly providing information about deposition and erosion processes at the bed (Gillette and Pitchford, 2004; Gillette et al., 2006). Another approach to investigate aeolian sediment transport is the use of fluorescent tracer sand, but it has only been used on unvegetated surfaces to date (Berg, 1983; Willetts and Rice, 1988; Cabrera and Alonso, 2009). More detailed information on deposition processes and wake development comes from wind tunnel experiments with model canopies of solid roughness elements (Sutton and McKenna Neuman, 2008a,b; Gillies et al., 2006). Various methods have been used to visualize bed-level flow and erosion patterns around solid roughness elements in wind tunnel experiments. Flow visualization techniques include ink and oil methods (Langston and Boyle, 1982; Bullard et al., 2000; Sutton and McKenna Neuman, 2008b) and a method based on the ammonia–manganese chloride reaction (Donat and Ruck, 1999). Sutton and McKenna Neuman (2008a) examined sediment transport initiation downwind of solid roughness elements by placing strips of sand across the wind tunnel floor and observing where they had eroded after exposing them to a wind event. However, results from experiments with solid roughness elements do not necessarily apply to live plant canopies (Walter et al., 2012). Since live plants are porous and flexible, they affect the wind flow and sediment flux differently than solid, rigid objects. Plants allow the wind and sediment particles to pass through them, and they have larger drag coefficients than solid objects, i.e. they can produce greater overall flow resistance because they have a larger surface subject to skin friction (Gillies et al., 2002; Gromke and Ruck, 2008). The study presented here differs from previous wind tunnel research in two respects. First, we used live vegetation instead of artificial roughness elements. Second, we used colored sand to visualize spatial patterns of sediment deposition within those vegetation canopies. The objective of this study was to identify and locate areas experiencing net deposition in three canopy densities of Perennial Ryegrass (Lolium perenne). Following a visual determination of net deposition patterns in different flow regimes, we estimate the fractional areas of sheltered and erodible ground surface within these grass canopies. K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 67 allowed firm rooting of the plants and provided a high waterretention capacity. To create a readily erodible surface, a 1 cm thick layer of quartz sand was spread over the planted soil. The grain size distribution of this sand is presented in Table 2. The third and fifth tray downstream were filled with red and blue resin-coated sand respectively, which was produced from the uncolored sand (Carlo Bernasconi AG, Switzerland, Fig. 2). Effects of the resin-coating on the erodibility of the sand grains, e.g. possible impacts on its grain size distribution or inter-particle electrostatic forces (Rasmussen et al., 2009), were not considered in this study. The frontal area index k was determined based on scaled frontal view images of individual plants. It was calculated as k = na/A, with n being the number of tussocks with a mean frontal area a occupying the ground area A (Table 1). At the time of the wind tunnel experiments, the tussocks had a mean height h of 10 cm (sd = 1, n = 20) and a mean frontal area a of 64 cm2 (sd = 18, n = 20). The time between experiments was at least 24 h, which allowed the plants to return to an upright position. During the 4 weeks of experimental work, the plants did not grow observably and they showed no visual evidence of sandblast injury. Vegetation cover Cv was defined as the proportion of the ground surface occupied by a vertical projection of the plants’ aerial parts (Greig-Smith, 1964 in Collins and Becker, 2001). It was determined based on scaled nadir images of whole plant trays (Fig. 3). Images were digitized and analyzed with the software ImageJ (Version 1.39u, http://rsb.info.nih.gov/ij/). 2. Material and methods 2.1. Wind tunnel facility The experiments were performed in the boundary layer wind tunnel of the WSL Institute for Snow and Avalanche Research SLF in Davos, Switzerland, located at 1650 m a.s.l. (Clifton et al., 2006). The wind tunnel operates in suction mode, with a fan downstream of the test section drawing air from outside through a honeycomb. It has a nominal cross-section of 1 m 1 m and a total length of 15 m. For the experiments described here, that length included a 2 m contraction section (4:1), a 5 m inlet duct with a smooth wooden floor, and an 8 m test section with an adjustable floor. For each experiment, eight wooden trays (98 99 cm) were installed in the test section of the wind tunnel, planted with a prescribed number of grass tussocks per unit ground surface (Fig. 2). The test section floor was lowered so that the surface matched the upwind inlet section, and the height of the wind tunnel roof was adjusted for each canopy density to provide a zero streamwise pressure gradient. 2.2. Vegetation Wind tunnel experiments were performed with three canopy densities of Perennial Ryegrass (Lolium perenne cv. Alligator, Alpine Garden Center Schutz Filisur, Switzerland), namely 5.25, 24.5 and 91 tussocks per square meter (Table 1). In the following discussion they are referred to as the low-, medium- and high-density canopies. The arrangement of the plants followed the same pattern of staggered rows in all experiments (Fig. 2). Lolium perenne was grown in a commercial horticulture nursery substrate (Topferde 140, Ricoter, Switzerland) in plant cultivation trays with integrated conical cells (20 ml). After seedlings reached a height of approximately 6 cm, they were removed from the cultivation trays with their entire root balls and planted in the eight wooden trays used for the wind tunnel experiments. The trays were filled with a 3:1 mixture of crushed limestone sand (grain sizes 0–3 mm, pH 8.4) and soil (grain sizes 0–2 mm, pH 7.5), which 2.3. Wind tunnel experiments The wind tunnel experiments discussed here were conducted as a component of a larger study of sediment mass fluxes in vegetation canopies described in more detail by Burri et al. (2011). The first experiment had a vegetation density of 91 plants per square meter. Subsequently, plants were removed from the trays to conduct experiments with area-averaged densities of 24.5 and 5.25 plants per square meter. After each experiment, the 1 cm quartz sand layer was renewed to provide consistent surface conditions, i.e. the eroded sand was replaced by new sand and the sand bed was flattened. wind tunnel side wall c wind a b c wind tunnel side wall 8m Fig. 2. Wind tunnel test section with eight aligned plant trays (98 99 cm), the third and fifth tray downstream being filled with red and blue quartz sand, respectively. Lowdensity canopy (top left), medium-density canopy (top middle), high-density canopy (top right), and schematic representation (bottom). a = upstream part of the plant tray, b = downstream part of the plant tray, c = lateral parts of the plant tray close to the side walls of the wind tunnel. (For interpretation of the references to color in this figure legend, the reader is referred to the web version of this article.) 68 K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 Table 1 Characteristics of the three canopy densities and experimental conditions. The air temperature [°C], relative air humidity [%] and matric potential of the sand [MPa] were determined as described in Burri et al. (2011). Planting density [plants/m2] Vegetation cover Cv [%] Frontal area index k Inter-plant spacing d [cm] Mean canopy height h [cm] Mean free stream velocity [m/s] Duration of erosive event [s] Air temperature [°C] Relative air humidity [%] Matric potential of the sand [MPa] Lowdensity Mediumdensity Highdensity 5.25 4 0.03 56 10 15.4 174 7.3 81.3 10.8 24.5 16 0.16 28 10 15.7 610 7.4 68.3 21.4 91 47 0.58 14 10 15.6 7224 9.9 54.5 25.1 Table 2 Grain size distribution of the quartz sand used in this study. Grain size class [mm] Percentage [%] >0.80 0.63–0.80 0.40–0.63 <0.40 4 54 40 2 Each canopy was subjected to an erosive wind event consisting of three phases. The wind tunnel motor was programmed to accelerate within 120 s to a free stream wind velocity of 15.5 m/s, this velocity was maintained for a set time and then the motor decelerated to 0 m/s within 100 s. Depending on the intensity of erosion in a given experiment, the duration of constant wind velocity was varied to yield a sufficient amount of erosion for recognizing patterns in the colored sand while not depleting the loose quartz sand on the beds. 2.4. Analysis of deposition patterns The analysis of deposition patterns described here is restricted to the trays containing red and blue sand, subsequently referred to as the red and blue sections (Fig. 2). Deposition in these trays was more obvious than in the six white trays because all the sand invading from upstream was of a different color than the sand with which the trays were originally filled. In contrast, the areas of sand deposition in the trays with white sand were less pronounced because the deposits contained both colored and white grains. There were 2 m of white sand upstream of the red section (Fig. 2). The source area upstream of the blue section included 3 m of white sand and 1 m of red sand (Fig. 2). Thus, all white sand grains that were found in the red section and all white and red sand grains in the blue section after the wind tunnel experiments represented deposits originating from upstream. To describe the locations of sediment deposits, we divided the sand bed into the following zones, according to their position related to a given tussock (Fig. 3): (a) the bleed flow zone, referring to the space within reach of the moving grass blades, projected on the ground, (b) the lateral zones, (c) the upstream area, and (d) the wake area. The elongated bare soil areas oriented in the direction of the wind are referred to as ‘streets’ (Gillette and Pitchford, 2004). Image sequences were taken through the perspex roof of the wind tunnel to evaluate the temporal development of deposition patterns during the experiments. In the low- and medium-density canopies, sediment flux was high and the deposited sediments were clearly visible. In the high-density canopy case, net sediment flux was low and it was difficult to detect the deposited sand grains in the images. In light of this, two different strategies were applied to analyze the deposition patterns in the different canopy densities. In the low- and medium-density cases a set of three consecutive images were selected to represent an initial, intermediate and final stage of each experiment (Figs. 4 and 5). The time intervals between the images were not recorded and put in relation to the duration of the experiment. Therefore, the image sequences only allow qualitative analysis of the temporal development of deposition patterns. The deposition patterns were intensified by increasing contrast, saturation and brightness of these images with Adobe Photoshop CS4. In the high-density canopy, the analysis of deposition patterns in these image sequences was supplemented with close-up images that were taken after the experiments (Fig. 6). The individual deposited grains were automatically selected based on their color, marked in black and enlarged by using Adobe Photoshop CS4. The automatic selection was then checked manually to ensure that the marked areas corresponded to deposited grains. In addition to describing the location of sand deposition relative to the plants and their flow characteristics, we estimated the fractions of the ground surface that were sheltered and potentially exposed to wind erosion in each experiment. We defined the sheltered portion (SP) as the fraction of the total surface that was Fig. 3. Schematic flow areas around a tussock: (a) bleed flow zone, (b) lateral zones, (c) upstream area, (d) wake area. The elongated bare soil areas that are oriented in the direction of the wind are referred to as ‘streets’ (dark shaded areas). K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 69 Fig. 4. Image sequences of sediment deposition patterns in the low-density canopy (inter-plant spacing = 56 cm). In the red section (A1–A3), white sand was invading from upstream. The dark spots at the top of image A3 are areas where erosion exposed the substrate underneath the red sand. In the blue section (B1–B3), red and white sand was invading from upstream. Wind direction: from left to right. (For interpretation of the references to color in this figure legend, the reader is referred to the web version of this article.) either covered by plants or experienced net deposition during the experiment. The remaining ground surface, which was potentially exposed to erosion, was defined as the erodible portion EPtot. EPtot is defined here relative to the total surface area, while EPnc, is defined in relation to the non-covered surface, i.e. the total surface area minus the area covered by grasses. The determination of SP and EPtot was made by visually discriminating between the colors of deposited sand and the sand with which the trays were filled originally. These discriminations were made while adhering to the following criteria and assumptions. Areas completely covered by upstream colored sand were attributed to SP, as they clearly experienced net deposition. Regions containing ripple patterns of alternating colors were attributed to EPtot, even if a substantial portion was covered by upstream colored sand. Sand in these ripples was assumed not to represent net deposition, as ripples migrate and are thus only temporary deposition sites (Fig. 7). Areas completely covered by sand with which the trays were filled originally were also attributed to EPtot, because these areas did not experi- Fig. 5. Image sequences of sediment deposition patterns in the medium-density canopy (inter-plant spacing = 28 cm). In the red section (A1–A3), white sand was invading from upstream. In the blue section (B1–B3), red and white sand was invading from upstream. Wind direction: from left to right. (For interpretation of the references to color in this figure legend, the reader is referred to the web version of this article.) 70 K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 Fig. 6. Sand grains deposited in the red and blue section of the high-density canopy (inter-plant spacing = 14 cm). The grains were automatically selected based on their color, marked in black and enlarged by using Adobe Photoshop CS4. Wind direction: from left to right. (For interpretation of the references to color in this figure legend, the reader is referred to the web version of this article.) Fig. 7. Visually determined proportion of sheltered surface SP in the red and blue section of the low-density canopy (top, inter-plant spacing = 56 cm) and the mediumdensity canopy (bottom, inter-plant spacing = 28 cm). Wind direction: from left to right. (For interpretation of the references to color in this figure legend, the reader is referred to the web version of this article.) ence net deposition. However, areas with unchanged color did not provide conclusive evidence for or against the following sediment transport scenarios: (i) neither erosion nor deposition, (ii) deposited grains originated in the same tray, (iii) more erosion than deposition (net erosion). The sheltered ground portions SP were manually selected and marked in the images representing the final stages of the experiments using Adobe Photoshop CS4 (Fig. 7). 3. Results 3.1. High-density canopy In the high-density canopy, the bleed flow zones of neighboring tussocks overlapped, covering almost the whole surface. Because net sediment flux was low, only isolated deposited sand grains K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 were found (Fig. 6). Such grains were distributed throughout the canopy, with particularly high concentrations in the bleed flow zones and lower concentrations in the plant interspaces. Within the bleed flow zones, the deposited grains were evenly distributed around the tussocks without pronounced accumulations on the upstream, downstream or lateral sides of the tussocks. 3.2. Medium-density canopy In the medium-density canopy, deposition occurred primarily in the bleed flow zones and wake areas (Figs. 3 and 5). In some parts of the canopy, deposition was also observed in the lateral zones. In general, the sand deposits in the wake areas were wedge-shaped in plan view and overlapped with the adjacent downstream tussocks. In the initial stage of the experiment (Fig. 5A1 and B1), deposition in the wake areas of the plants occurred primarily at the upstream ends of the red and blue sections, with less intense deposition occurring in the downstream areas of those trays (Figs. 2 and 5). In the intermediate stage (Fig. 5A2 and B2) deposition became more intense over the whole sections and the deposition areas in the wakes of the tussocks expanded laterally. During this intermediate stage of the experiment, rudimentary ripples evolved in the streets between the tussocks. In the final stage of the experiment (Fig. 5A3 and B3) the deposition areas increased in width, and expanded into the open streets between the tussocks in the central areas of the trays. Along the outside edges of the plant trays, i.e. between the wind tunnel side walls and the second plant rows from the edges, there was less deposition than in the center of the trays during all three experiment stages (Figs. 2 and 5). The deposition areas were generally larger in the blue section than in the red section (e.g. Fig. 5A3 and B3). The sheltered ground surface SP equaled 33% in the red section and 62% in the blue section (Fig. 7 and Table 3). The erodible portion EPnc accounted for 77% of the surface not covered by grasses in the red section and 44% in the blue section (Table 3). 3.3. Low-density canopy In the low-density canopy, discrete deposition patterns could be observed in all four aerodynamic zones around the tussocks: (a) the bleed flow zone, (b) the lateral zones, (c) the upstream area, and (d) the wake area (Figs. 3 and 4). In contrast to the mediumdensity canopy, the wake deposits did not overlap with the adjacent downstream tussocks. In the initial stages of the experiments (Fig. 4A1 and B1) deposition was found in all four zones mentioned above. The deposits in the wake areas were wedge-shaped in plan view tapering to indeterminate downwind ends during the initial stage. In the intermediate and final stages of the experiments (Fig. 4A2–A3 and B2–B3) the wake deposits expanded laterally and a clearly discernible ripple pattern evolved in the streets between the tussocks. The Table 3 Portions of sheltered surface SP, and erodible surface in relation to the total surface EPtot and in relation to the surface which was not covered by grasses EPnc. Low-density canopy Medium-density canopy Red section Blue section Red section Blue section Grass-covered area [%] Not-covered portion [%] Sheltered portion SP [%] Erodible portion EPtot [%] Erodible portion EPnc [%] 3 97 24 76 78 5 95 36 64 67 13 87 33 67 77 13 87 62 38 44 71 downstream ends of the deposits in the wakes of the tussocks became more pronounced due to the development of ripples approximately 10 cm upwind of the adjacent downstream tussocks. Analogous to the medium-density canopy case, there was more deposition in the blue than in the red tray, and more deposition in the center than in the areas close to the side walls of the wind tunnel in all three stages (Figs. 2 and 4). The sheltered ground surface SP amounted to 24% of the red section and 36% of the blue section (Fig. 7 and Table 3). The erodible area EPnc totaled 78% of the noncovered red section and 67% of the blue section. 4. Discussion The spatial patterns of colored sand found in this study demonstrate that multiple processes control aeolian sediment entrainment and deposition on a vegetated surface. One important process was sediment deposition in the sheltered wake areas of the tussocks. In the low- and medium-density canopies, such wake deposits were the primary regions of sediment deposition (Figs. 4 and 5). The size of these wake deposits in relation to the plant spacing indicates that the two canopy densities represented an isolated roughness flow and a wake-interference flow regime, respectively. In the low-density canopy, the wake deposits did not intercept the adjacent downwind tussocks, while in the mediumdensity canopy they overlapped with the downstream vegetation. In the high-density canopy, the lack of wake deposits and the even distribution of deposited sand grains within the bleed flow zones (Fig. 6) suggests a skimming flow regime in which the whole surface was sheltered, with no individual wake areas. In the immediate vicinity of the grass tussocks, i.e. within reach of the moving grass blades, windborne particles were deposited when they hit plant surfaces, lost momentum and fell. This process caused the sediment accumulations in the bleed flow zones around the tussocks. It was probably also responsible for deposits in the areas upstream of the tussocks in the low-density canopy, visible in the last pictures of the image sequences (Fig. 4A3 and B3). Some of the windborne particles in the sand cloud approaching the tussocks may have rebounded from the grass blades and come to settle in front of the tussocks (Bagnold, 1941). In the high- and medium-density canopies the wake areas overlapped with the downstream tussocks so deposits upstream of a given tussock could not be identified as up-wind deposits or deposits in the wake of the upstream tussock (Figs. 5 and 6). Walter et al. (2012) measured the bed level shear stress distribution around tussocks of L. perenne and solid cylinders. Compared to a situation without a roughness element, they found increased shear stress to the sides of both the grass tussock and the cylinder, with a higher peak shear stress adjacent to the cylinder than the grass tussock. This reflects the fact that flow acceleration around a solid obstacle is higher than around porous obstacles (Bowker et al., 2008). Hence, more sediment entrainment is to be expected around a solid, rigid obstacle than around a porous, flexible obstacle. Leenders et al. (2007) observed in their field study that sediment accumulated upstream and laterally of a single shrub, whereas sediment scouring occurred around a single tree trunk. In the present study we found sediment deposits immediately to the sides of the tussocks, indicating that the shear stress acting on the ground was below the threshold for sediment entrainment. This is most evident in Fig. 4A3, where these lateral deposits are clearly larger than the bleed flow zones. In the low-density canopy, the tussocks in the lateral parts of the plant trays close to the side walls of the wind tunnel produced asymmetric deposition without lateral deposition on their outer sides (Figs. 2 and 4). This phenomenon is probably caused by enhanced turbulences close to the side walls of the wind tunnel 72 K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 due to flow recirculation and corner vortices. Further evidence for enhanced erosion near the side walls of the wind tunnel was found in the medium-density canopy, where erosion was particularly intense in the streets adjacent to the side walls, i.e. between the outermost and second outermost plant rows (Fig. 5). The influence of the tunnel side-walls died out within two plant-spacings from the edge, and was not observed in the central portion of the tunnel on which the results presented here are based. The areas of net deposition were generally smaller in the red section than in the blue section. Since the red section was closer to the upwind beginning of the test section, it had a smaller sediment fetch than the blue section. Aeolian sediment flux tends to increase up to saturation with the distance over which the erosion occurs (the ‘‘fetch effect’’, Gillette et al., 1996). Additionally, the characteristic flow regimes of the three canopy densities required some distance to fully develop because the entering air encountered a change in surface roughness when it passed from the smooth inlet duct of the wind tunnel to the planted test section. Therefore, the near ground wind velocities were probably highest and most efficient in entraining sediment at the beginning of the test section, leading to higher erosion rates near the beginning of the test section than towards the downwind end. The changing erosive capacity of the flow along the test section offered an opportunity to study the development of deposition patterns under different erosion intensities. In the course of an erosive event, the sand deposit in the wake of an obstacle gradually increases in areal cover and height until its slopes stand at the limiting angle of repose (Bagnold, 1941). Once this angle is reached, any additional sand slips down the slopes and leaves the sheltered wake area, so it can be entrained by the stronger wind outside the wake. In the blue section where the erosive capacity of the wind flow was lower than in the red section this process enabled the deposition areas to spread into the open streets between the tussocks (Figs. 4B3 and 5B3). In the red section, in contrast, there was only minor deposition in the streets (Figs. 4A3 and 5A3). Accordingly, the erodible surface areas EPtot were higher in the red section than in the blue section (Table 3). In spite of these differences between the red and blue sections, EPtot was in any case substantially smaller than the ground surface that was not covered by grasses (Table 3). Many wind erosion models for vegetated surfaces, however, approximate the erodible area as the total surface area not covered by non-erodible elements (e.g. Marticorena and Bergametti, 1995). The present study supports the findings of Chappell et al. (2010) who showed that this approximation can lead to substantial over-estimates of aeolian sediment fluxes. Chappell et al. (2010) developed a model based on angular reflectance and estimated the erodible portion for different arrays of solid hemispheres. For an array with a vegetation cover Cv of 5% and a frontal area index k of 0.04, they estimated EPnc to be 83% of the non-covered surface. The visually determined EPnc in our low-density canopy, which had a similar Cv and k (Table 1), was 78% of the non-covered surface in the red section and 67% in the blue section (Table 3), demonstrating that the erodible surface fraction in canopies of flexible, porous grass tussocks is even smaller than in comparable arrays of solid roughness elements. For future studies it would be interesting to investigate in more detail the performance of models that approximate the erodible area as the uncovered area, e.g. by validating them with laboratory wind tunnel measurements. The present wind tunnel study was distinct from previous research in that colored sand was used to examine erosion and deposition processes in live plant canopies. Colored sand proved to be useful for this purpose, as it produced high-resolution bed-level data for the whole ground surface under investigation. This is of particular importance when dealing with live plants, as they are non-uniform complex structures that interact with the wind in more variable ways than solid artificial roughness elements. The application of colored sand is a relatively easy and low-cost method to study aeolian sediment transport in vegetation canopies, and it may therefore be useful not only for wind tunnel studies, but also for larger scale field studies. The sensitivity of the method, i.e. the detectability of colored sand deposits depends on several factors, such as the scale of the images, the applied image analysis procedure and the characteristics of the underlying surface. Previous field studies have used fluorescent tracer sand on unvegetated surfaces and transferred deposits via sticky tapes to the laboratory for analysis under ultraviolet light (Berg, 1983; Cabrera and Alonso, 2009). The use of colored resin-coated sand, in contrast, may be suitable to collect on-site data, by taking high-resolution close-up images directly in the field. 5. Conclusions The present study describes wind tunnel experiments with live grass tussocks. The spatial patterns of redistributed colored sand within a high-, medium- and low-density canopy indicated three distinct aerodynamic flow regimes, i.e. a skimming flow, a wakeinterference flow and an isolated roughness flow. Previous research on the factors determining which air flow regime a canopy will experience has been based on experiments with solid roughness elements (Morris, 1955; Lee and Soliman, 1977; Wolfe and Nickling, 1993). In the low and medium-density canopies, sediment deposits were primarily found in the wake areas downstream of the tussocks. While they overlapped with the adjacent downstream tussocks in the medium-density canopy, they did not in the lowdensity canopy. In the high-density canopy, very few sand grains were entrained by the wind, and the deposited grains were evenly distributed within the disturbed zones surrounding the grass tussocks. We found that the fraction of the sand surface which effectively experienced erosion (EPnc) was substantially smaller than the area which was not covered by grasses in three vegetation canopy densities associated with different flow regimes. The three experiments conducted in this study do not allow us to estimate the proportionality factor between EPnc and vegetation density. Further studies are needed to establish this relationship under various conditions. While the values of EPnc and the boundary conditions for each flow regime in this experimental study may not be exactly replicated by field studies under natural wind conditions, our general findings and trends should be representative. Acknowledgments This study and the main author were financially supported by the Velux Foundation. Additional funding was provided by the Swiss National Science Foundation and the Vontobel-Foundation. Sensirion AG kindly provided relative humidity sensors and technical support. The Alpine Garden Center Schutz Filisur shared their knowledge with us and provided access to their plant nursery facilities. We wish to thank Dr. Costantino Manes for his help in planning and designing this study and Dr. Martin Schneebeli for his assistance with the software ImageJ. We would also like to thank Silvio Burger, Hans Herranhof, Andreas Moser, Christian Simeon and Andreas Tröger for their hard work in the wind tunnel and for manufacturing several experimental devices. We wish to acknowledge in particular the constructive and very helpful suggestions of Prof. Ottmar Holdenrieder, Prof. Michael Lehning and Benjamin Walter that led to several important refinements of the manuscript. K. Suter-Burri et al. / Aeolian Research 8 (2013) 65–73 References Bagnold, R.A., 1941. 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